2021 in paleomalacology explained

This list of fossil molluscs described in 2021 is a list of new taxa of fossil molluscs that were described during the year 2021, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2021.

Ammonites

New taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages
Abichites infirmis[1] Sp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Abichites ovalisSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Agathiceras iranicum[2] Sp. novValidKorn & Ghaderi in Korn et al.Carboniferous-Permian boundary
Alienisphinctes[3] Gen. et sp. novValidÉnay & MangoldLate Jurassic (Kimmeridgian)Jubaila FormationA member of the family Ataxioceratidae. The type species is A. cavelieri.
Amaltheus orientalis[4] Sp. novValidNakada et al.Early Jurassic (Pliensbachian)Kuruma Group
Arasella falcataSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast FormationAn ammonite of uncertain placement, probably a member of the family Xenodiscidae.
Arcestes lawsi[5] Sp. novValidTaylor, Guex & LucasLate Triassic (Rhaetian)Gabbs Formation
Arnioceras ritterbushi[6] Sp. novValidTaylor & GuexEarly Jurassic (Sinemurian)Sunrise Formation
Arnioceras sparsumSp. novValidTaylor & GuexEarly Jurassic (Sinemurian)Sunrise Formation
Artyomites[7] Gen. et sp. novValidSmyshlyaeva & ZakharovEarly Triassic (Olenekian)A member of the family Palaeophyllitidae. Genus includes new species A. antiquus.
Aulatornoceras frenklerae[8] Sp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Aulatornoceras steinhauseniSp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Aulatornoceras ventrosulcatumSp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Boucaulticeras hawthornensisSp. novValidTaylor & GuexEarly Jurassic (Sinemurian)Sunrise Formation
A member of the family Schlotheimiidae.
Bullatimorphites szontaghi[9] Sp. novValidGalácz
Caloceras avus[10] Sp. novValidKmentEarly Jurassic (Hettangian)
Catrisimoceras[11] Gen. et comb. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and possibly to the subfamily Lytogyroceratinae. The type species is "Simoceras" catrianum Zittel (1870).
Cheltonia howarthi[12] Sp. novValidSimms & EdmundsEarly Jurassic
Cibolaites petraschecki[13] Sp. novValidWilmsen et al.Late Cretaceous (Turonian)Brießnitz Formation
ClivotirolitesGen. et 2 sp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast FormationA member of the family Dzhulfitidae. The type species is C. decoratus; genus also includes C. petilus.
Crassotornoceras hetzeneggeriSp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Crassotornoceras nudumSp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Crioceratites claveli[14] Sp. novIn pressBert et al.Early Cretaceous (Hauterivian)Vocontian Basin
Didymoceras morozumii[15] Sp. novValidMisaki & TsujinoLate Cretaceous (Campanian)Hiketa Formation
Dimorphodiscus[16] Gen. et sp. et comb. novValidVermeulen et al.Early Cretaceous (Barremian)
Crimea
A member of the family Holcodiscidae. The type species is D. lerisae; genus also includes D. uhligi (Karakash, 1907), D. sayni (Joleaud, 1912) and "Metahoplites" postnodosum Vermeulen et al. (2017).
Discoscaphites mullinaxorum[17] Sp. novValidWitts et al.Late Cretaceous (Maastrichtian)Corsicana Formation
Domanikoceras martenbergense[18] Sp. novValidKornDevonian (Frasnian)A member of the family Tornoceratidae.
Dzhulfites brevisellatusSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Eoasianites baghevangensisSp. novValidKorn & Ghaderi in Korn et al.Carboniferous-Permian boundary
EoermocerasGen. et 2 sp. novValidÉnay & MangoldMiddle Jurassic (Bajocian)Dhruma FormationA member of the family Stephanoceratidae. The type species is E. glabrum; genus also includes new species E. angulatidomus.
Eolytoceras rohrbachensisSp. novValidKmentEarly Jurassic (Hettangian)
EsfahanitesGen. et sp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast FormationA member of the family Dzhulfitidae. The type species is E. armatus.
Eutomoceras paladeji[19] Sp. novValidTongtherm, Nabhitabhata & TuanapayaTriassicChaiburi FormationA member of the family Ceratitidae.
Eutomoceras wantanaeSp. novValidTongtherm, Nabhitabhata & TuanapayaTriassicChaiburi FormationA member of the family Ceratitidae.
Evopharciceras[20] Gen. et sp. et comb. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein Formation
A member of Agoniatitida belonging to the family Pharciceratidae. The type species is E. formosum; genus also includes "Pharciceras" evolvens Bockwinkel, Becker & Ebbighausen (2009).
Extropharciceras metallicumSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
Galfettites orientalisSp. novValidSmyshlyaeva & ZakharovEarly Triassic (Olenekian)A member of the family Galfettitidae.
Glyptophiceras mirumSp. novValidSmyshlyaeva & ZakharovEarly Triassic (Olenekian)A member of the family Xenoceltitidae.
Gonioclymenia junggarensis[21] Sp. novIn pressZong, Becker & MaDevonian (Famennian)Hongguleleng Formation
Hadrothisbites hanwangensis[22] Sp. novIn pressMietto et al.Late Triassic (Carnian)Ma'antang Formation
Holcodiscus pseudocaillaudiSp. novValidVermeulen et al.Early Cretaceous (Barremian)A member of the family Holcodiscidae.
Hulenites kitamuraiSp. novIn pressMatsukawaEarly Cretaceous (Aptian)Todai Formation
Inaptychoceras[23] Gen. et sp. novIn pressMatsukawaEarly Cretaceous (Aptian)Todai FormationGenus includes new species I. koikei.
Kislovodskoceras[24] Gen. et sp. novValidPolkovoyEarly Cretaceous (Aptian)A member of the family Desmoceratidae. The type species is K. kislovodskense.
Kurnubiella fallaxSp. novValidÉnay & MangoldMiddle Jurassic (Callovian)Upper Dhruma FormationA member of the family Pachyceratidae.
LaevisimocerasGen. et sp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is L. inornatum.
Linguatornoceras asbeckenseSp. novValidKornDevonian (Frasnian)A member of the family Tornoceratidae.
Lunupharciceras kochiSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
LutitesGen. et 5 sp. et comb. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast FormationA member of the family Dzhulfitidae. The type species is L. lyriformis; genus also includes new species L. alius, L. paucis, L. plicatus and L. profundus, as well as Alibashites stepanovi Korn & Ghaderi in Korn et al. (2015).
Lytogyroceras tuberculatumSp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Lytogyroceratinae.
Maenioceras ornatumSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Maenioceratidae.
Nejdia duartei[25] Sp. novValidGoy & Comas-RengifoEarly Jurassic (Toarcian)A member of the family Hildoceratidae.
Obtusicostites gigasSp. novValidÉnay & MangoldMiddle Jurassic (Callovian)Upper Dhruma FormationA member of the family Perisphinctidae.
Ophimullericeras[26] Gen. et sp. novValidJenks et al.Early TriassicDinwoody Formation
A member of Ceratitida belonging to the group Meekocerataceae and the family Mullericeratidae. The type species is O. paullae.
Pachyerymnoceras arabicumSp. novValidÉnay & MangoldMiddle Jurassic (Callovian)Tuwaiq Mountain Limestone FormationA member of the family Pachyceratidae.
Pachyerymnoceras magnumSp. novValidÉnay & MangoldMiddle Jurassic (Callovian)Tuwaiq Mountain Limestone FormationA member of the family Pachyceratidae.
Palaeophyllites mishkiniSp. novValidSmyshlyaeva & ZakharovEarly Triassic (Olenekian)A member of the family Palaeophyllitidae.
Paracadoceras vasily[27] Sp. novValidGulyaev in Gulyaev & IppolitovMiddle Jurassic (Callovian)
Paracochloceras nunminensisSp. novValidTaylor, Guex & LucasLate Triassic (Rhaetian)Gabbs Formation
Parahoplites inaensisSp. novIn pressMatsukawaEarly Cretaceous (Aptian)Todai Formation
ParasimocerasGen. et 2 sp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. venturii; genus also includes P. pseudostenonis.
Paratirolites aduncusSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Paratirolites baghukensisSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Paratirolites lanceolobatusSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Paratirolites robustusSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Paratirolites rubensSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast Formation
Peripleurites gabbensisSp. novValidTaylor, Guex & LucasLate Triassic (Rhaetian)Gabbs Formation
A member of Choristocerataceae belonging to the family Rhabdoceratidae.
Pharciceras beyrichiSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
Pharciceras ferrumSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
Pharciceras kruegeriSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
Placites heggiSp. novValidTaylor, Guex & LucasLate Triassic (Rhaetian)Gabbs Formation
A member of the family Gymnitidae.
PlanisimocerasGen. et sp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. costotuberculatum.
Pluripharciceras ahlburgiSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
Pronjaites yolkinensis[28] Sp. novValidMittaEarly Cretaceous
PseudogalfettitesGen. et sp. novValidSmyshlyaeva & ZakharovEarly Triassic (Olenekian)A member of the family Galfettitidae. Genus includes new species P. sakhnoi.
Psiloceras pseudocostosumSp. novValidKmentEarly Jurassic (Hettangian)
Retrotornoceras juxiSp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Rhacophyllites mulleriSp. novValidTaylor, Guex & LucasLate Triassic (Rhaetian)Gabbs Formation
Rhacophyllites volcanoensisSp. novValidTaylor, Guex & LucasLate Triassic (Rhaetian)Gabbs Formation
Rollierites biplicatum[29] Sp. novValidJain & MazurMiddle Jurassic (Callovian)A member of the family Pachyceratidae.
Sandbergeroceras archiaciSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Triainoceratidae.
SaudicostitesGen. et sp. novValidÉnay & MangoldMiddle Jurassic (Bathonian)Dhruma FormationA member of Perisphinctoidea of uncertain phylogenetic placement. The type species is S. arabicus.
Shevyrevites corrugatusSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast FormationA member of the family Xenodiscidae.
Sigaloceras fundatorSp. novValidGulyaev in Gulyaev & IppolitovMiddle Jurassic (Callovian)
Sigaloceras kiseleviSp. novValidGulyaev in Gulyaev & IppolitovMiddle Jurassic (Callovian)
SimocosticerasGen. et sp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. biornatum.
SinotropitesGen. et sp. novIn pressMietto et al.Late Triassic (Carnian)Ma'antang FormationGenus includes new species S. sichuanensis.
Sonninia (Euhoploceras) crassicostaeSp. novValidÉnay & MangoldMiddle Jurassic (Bajocian)Lower Dhruma FormationA member of the family Sonniniidae.
Sonninia (Euhoploceras) turbatorSp. novValidÉnay & MangoldMiddle Jurassic (Bajocian)Lower Dhruma FormationA member of the family Sonniniidae.
Sonoraceras[30] Gen. et sp. novIn pressSamaniego-Pesqueira, Moreno-Bedmar & Álvarez-SánchezEarly Cretaceous (Aptian)Genus includes new species S. tepachensis.
Stenopharciceras lotziSp. novValidKorn & BockwinkelDevonian (Givetian)Roteisenstein FormationA member of Agoniatitida belonging to the family Pharciceratidae.
Stoliczkaia (Lamnayella) duboisi[31] Sp. novValidKennedy & LatilLate Cretaceous (Cenomanian)A member of the family Lyelliceratidae.
Stoyanowites parallelusSp. novValidKorn & Hairapetian in Korn et al.Permian (Changhsingian)Hambast FormationA member of the family Dzhulfitidae.
Strenoceras djangurense[32] Sp. novValidMittaMiddle Jurassic (Bajocian)
SulcicostitesGen. et sp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. costatus.
Timorites dendroides[33] Sp. novValidKorn & Ghaderi in Tabrizi et al.Permian (Wuchiapingian)Julfa FormationA member of the family Cyclolobidae.
Tornoceras aequilobumSp. novValidSöte et al.Devonian (Frasnian)Sand FormationA member of the family Tornoceratidae.
Tornoceras fundibulumSp. novValidKornDevonian (Famennian)A member of the family Tornoceratidae.
Tornoceras occasus[34] Sp. novValidKornLate DevonianA member of the family Tornoceratidae.
Tornoceras ortusSp. novValidKornLate DevonianA member of the family Tornoceratidae.
Tornoceras vretterenseSp. novValidKornDevonian (Givetian)A member of the family Tornoceratidae.
Tropaeum evolutum[35] Sp. novIn pressLehmann & BaylissEarly Cretaceous (Aptian)Hythe Formation
Tropidoceras komarovi[36] Sp. novValidZaitsevEarly JurassicCrimean Peninsula
Vicininodiceras dalriatenseSp. novValidSimms & EdmundsEarly Jurassic
Volanoceras cuneospinatumSp. novValidNannarone & BilottaLate Jurassic (Tithonian)A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Volanoceratinae.
Waehneroceras morawetziSp. novValidKmentEarly Jurassic (Hettangian)
Wordieoceras mullenaeSp. novValidJenks et al.Early TriassicDinwoody Formation
A member of Ceratitida belonging to the group Meekocerataceae and the family Ophiceratidae.

Research

Other cephalopods

New taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages
Acrocoelites ippolitae[52] Sp. novValidWeis in Weis et al.Middle Jurassic (Aalenian)A belemnite.
Anglonautilus hewaidyi[53] Sp. novIn pressMoneer, Bazeen & KassabLate Cretaceous (Maastrichtian)A nautiloid cephalopod.
Angulithes monqarensisSp. novIn pressMoneer, Bazeen & KassabLate Cretaceous (Maastrichtian)A nautiloid cephalopod.
Arionoceras tokushimaense[54] Sp. novValidNikoSilurian (Ludlow)Suberidani Group
Bactroceras fluvii[55] Sp. novValidKröger & PohleOrdovician (Dapingian)Valhallfonna Formation
Boreopeltis ifrimae[56] Sp. novIn pressFuchs & StinnesbeckLate Cretaceous (Turonian)
Buttsoceras buldrebreenenseSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of the family Troedssonellidae.
Cycloplectoceras hinlopenseSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of the family Tarphyceratidae.
Cyclostomiceras profilstrandenseSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of the family Cyclostomiceratidae.
Deltoceras belugaSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of the family Cyclostomiceratidae.
Eosomichelinoceras borealisSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of the family Baltoceratidae.
Eosomichelinoceras submedius[57] Sp. novValidEvans et al.Ordovician (Darriwilian)Pelmis FormationA member of the family Baltoceratidae.
EthanocerasGen. et sp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA relative of Rioceras. The type species is E. solitudines.
Gzheloceras aisenvergi[58] Sp. novValidDernovCarboniferous (Serpukhovian)Samara FormationA member of the family Tainoceratidae.
Hemichoanella occultaSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna Formation
HinlopocerasGen. et 2 sp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA relative of Rioceras. The type species is H. tempestatis; genus also includes H. venti.
Jovellania praetermissa[59] Sp. novValidBrauckmann & KochDevonian (Givetian)Massenkalk Formation
Knightoceras extorrisSp. novValidDernovCarboniferous (Pennsylvanian)A member of the family Koninckioceratidae.
Lawrenceoceras ebenusSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna Formation
Lawrenceoceras larusSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna Formation
Litoceras profilbekkenenseSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna Formation
Mojsvaroceras gianii[60] Sp. novValidPieroni & PrinothMiddle Triassic (Anisian)A nautiloid.
NyfrieslandocerasGen. et sp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is N. bassleroceroides.
OlenidslettocerasGen. et sp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is O. farmi.
Paracenoceras waageni[61] Sp. novValidHalderMiddle Jurassic (Callovian)Chari FormationA member of the family Paracenoceratidae.
Planetoceras yefimenkoiSp. novValidDernovCarboniferous (Bashkirian)Mospinka FormationA member of the family Koninckioceratidae.
Proterocameroceras valhallfonnenseSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna Formation
Protocycloceras minorSp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA member of the family Troedssonellidae.
Rhynchorthoceras yizanense[62] Sp. novValidFang in Fang et al.OrdovicianA member of Lituitida.
Sinoceras complexumSp. novValidFang in Fang et al.OrdovicianA member of Lituitida.
SorosocerasGen. et sp. novValidEvans et al.Ordovician (Darriwilian)Pelmis FormationA member of the family Geisonoceratidae. The type species is S. castellum.
SvalbardocerasGen. et 2 sp. novValidKröger & PohleOrdovician (Floian)Valhallfonna FormationA relative of Rioceras. The type species is S. sterna; genus also includes S. skua.
Tyrioceras longicameratumSp. novValidFang in Fang et al.OrdovicianA member of Lituitida.
Xiaohenautilus mulleniSp. novValidJenks et al.Early TriassicDinwoody Formation
A member of Nautilida belonging to the group Trigonoceratoidea and the family Grypoceratidae.

Research

Gastropods

New taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages
Acirsa ambrosii[72] Sp. novValidBrunetti & CrestiPlioceneA species of Acirsa.
Acirsa (Notacirsa) texanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Acirsa.
Acteon pseudotypicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Acteon.
Acutitomaria[73] Gen. et sp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida. Genus includes new species A. kustatscherae.
Alatostrombus[74] Gen. et sp. et comb. novValidRavenMiocene
A member of the family Strombidae. Genus includes new species A. acutalatus, as well as "Strombus" sedanensis Martin (1899).
Alvania absuturalis[75] Sp. novValidChirli & ForliPlioceneA species of Alvania.
Alvania ameliaeSp. novValidChirli & ForliPleistoceneA species of Alvania.
Alvania benestarensisSp. novValidChirli & ForliMioceneA species of Alvania.
Alvania cerreti[76] Sp. novValidGardella et al.PlioceneA species of Alvania.
Alvania pluricosticillataSp. novDisputedChirli & ForliPlio-PleistoceneOriginally described as a species of Alvania. Argued to be a junior synonym of the extant taxon Crisilla gagliniae by Amati & Oliverio (2024).[77]
Amaea (Confusiscala) paleocaenicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Ammonicera danieli[78] Sp. novValidChernyshev & GoedertLate Eocene/early OligoceneGries Ranch Formation
A species of Ammonicera.
Ammonicera rolaniSp. novValidChernyshev & GoedertLate Eocene/early OligoceneGries Ranch Formation
A species of Ammonicera.
Ampezzogyra angulata[79] Sp. novValidNützel & Hausmann in Hausmann et al.Late Triassic (Carnian)San Cassiano FormationA member of the family Stuoraxidae.
Amphidromus ubaldii[80] Sp. novValidDharmaLate PlioceneA species of Amphidromus.
AmplitomariaGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" spuria Münster (1841).
Anacithara akisi[81] Sp. novValidKolokotronisPlioceneA species of Anacithara.
Anceps siminescuiNom. novValidHarzhauserMiddle-late MioceneA Trochidae member;
a replacement name for Trochus semistriatus Siminescu & Barbu (1940).
AngulatellaGen. et sp. novValidNützel & Hausmann in Hausmann et al.Late Triassic (Carnian)San Cassiano FormationA member of the family Prostyliferidae. The type species is A. bizzarinii.
Antithala[82] Gen. et comb. et 2 sp. novValidHarzhauser & LandauMioceneDej Formation



A member of the family Costellariidae. The type species is "Mitra" sturi Hoernes & Auinger (1880); genus also includes "Mitra" neugeboreni Hoernes & Auinger (1880), as well as new species A. claviformis and A. filipescui.
Apiocypraea cerretensis[83] Sp. novValidFehse et al.Pliocene (Zanclean)A member of the family Eocypraeidae.
Aplus abessensis[84] Sp. novValidLozouetOligocene (Chattian)A species of Aplus.
Aplus grandiculusSp. novValidLozouetOligocene (Chattian)A species of Aplus.
Aplus littoralisSp. novValidLozouetOligocene (Chattian)A species of Aplus.
Aplus pseudoaquitanensisSp. novValidLozouetOligocene (Chattian)A species of Aplus.
Aquilofusus? antiquaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Ascheria canni[85] Sp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A member of the family Hokkaidoconchidae.
Asiaharpa[86] Gen. et sp. novValidRavenMioceneA member of the family Harpidae. Genus includes new species A. sarawakiana.
Attenuata? danicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Possibly a species of Attenuata.
AuingeriaGen. et comb. novValidHarzhauser & LandauMioceneBaden Formation


A member of the family Columbellidae. The type species is "Columbella" bellardii Hörnes (1852); genus also includes "Columbella (Anachis)" austriaca Hoernes & Auinger (1880) and "Pyrene (Anachis) bellardii" grussbachensis Csepreghy-Meznerics (1969; raised to the rank of a separate species Auingeria grussbachensis).
Axonella[87] Gen. et comb. et sp. novValidPacaud, Lhomme & RenaudEocene (Ypresian)
A member of the family Hemisinidae. The type species is "Cerithium" palense Rouault (1850); genus also includes "Potamides" granosus Doncieux (1908), as well as new species A. falsiacum.
Babylonia chattica[88] Sp. novValidLozouetOligocene (Chattian)A species of Babylonia.
Babylonia edigittenbergeri[89] Sp. novValidRavenMioceneMiri FormationA species of Babylonia.
Babylonia sarawakianaSp. novValidRavenMioceneMiri FormationA species of Babylonia.
Bandellina compactaSp. novValidNützel & Hausmann in Hausmann et al.Late Triassic (Carnian)San Cassiano FormationA member of the family Cornirostridae.
Barycypraea abbasi[90] Sp. novValidDovesi & ParsonsMiocene (Serravallian)Nyalindung FormationA species of Barycypraea.
Barycypraea alessandrodovesiSp. novValidDovesi & ParsonsMiocene (Serravallian)Nyalindung FormationA species of Barycypraea.
Bela pediensis[91] Sp. novValidKolokotronisPlioceneA species of Bela.
BellacolumbellaGen. et comb. novValidHarzhauser & LandauMioceneBaden Formation


A member of the family Columbellidae. The type species is "Columbella (Nitidella)" karreri Hoernes & Auinger (1880); genus also includes "Columbella (Nitidella)" embryonalis Boettger (1902) and "Columbella (Nitidella)" katharinae Hoernes & Auinger (1880).
BellardithalaNom. et 4 sp. novValidHarzhauser & LandauMiocene and PlioceneBaden Formation
Dej Formation






?
A member of the family Costellariidae; a replacement name for Micromitra Bellardi (1888). The type species is "Voluta" obsoleta Brocchi (1814); genus also includes new species B. kovaci, B. baluki, B. fedosovi and B. dacica.
BieleraxisGen. et 2 sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species B. discoidea and B. altis.
Bithynia disregularis[92] Sp. novIn pressLi et al.OligoceneYehucheng FormationA species of Bithynia.
Bithynia obliquusSp. novIn pressLi et al.OligoceneYehucheng FormationA species of Bithynia.
Bithynia parainclinimargaSp. novIn pressLi et al.OligoceneYehucheng FormationA species of Bithynia.
Bithynia paramonolithicSp. novIn pressLi et al.OligoceneYehucheng FormationA species of Bithynia.
Bittium prekoeneniSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Bittium.
Bulimulus frenguellii[93] Sp. novValidCabrera et al.Late CretaceousQueguay FormationA species of Bulimulus.
Burminella[94] Gen. et sp. novIn pressBalashov, Perkovsky & VasilenkoLate Cretaceous (Cenomanian)Burmese amberA land snail possessing morphological traits intermediate between members of the family Pupinidae and other members of Cyclophoroidea. Genus includes new species B. artiukhini.
Bursa transeuntis[95] Sp. novValidForli & SmriglioMiocene and Pliocene

A species of Bursa.
Caecum texanumSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Caecum.
Califrapana[96] Gen. et comb. novValidPowell & HouartLate Oligocene to early MioceneVaqueros Formation

A member of the family Muricidae. The type species is "Purpura" vaquerosensis Arnold (1907).
Calyptraphorus velususSp. novValidGarviePaleocene (Danian)Kincaid Formation
Camponaxis bandeli[97] Sp. novValidPieroni, Monari & ToddLate Triassic (Carnian)San Cassiano FormationA Tofanellidae member
Campylacrum texanumSp. novValidGarviePaleocene (Danian)Kincaid Formation
Canariella (Majorata) gerti[98] Sp. novValidGrohQuaternaryA species of Canariella.
CancellotomariaGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" subcancellata d'Orbigny (1850).
Cancilla nanostriatula[99] Sp. novValidHarzhauser & LandauMioceneBaden Formation

A species of Cancilla.
Cancilla wagreichiSp. novValidHarzhauser & LandauMioceneDej FormationA species of Cancilla.
Capulus gelusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Capulus.
Cassis kalmani[100] Sp. novValidVicián & KovácsEocene (Lutetian)Csernye FormationA species of Cassis.
Cassis rasae[101] Sp. novValidPetuch & BerschauerPleistocene
A species of Cassis.
Cassis viliusiSp. novValidPetuch & BerschauerPleistocene
A species of Cassis.
CenorhytisGen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species C. groesbecki.
Ceratopea? moridunensis[102] Sp. novValidEbbestad & CopeOrdovician (Floian)A member of the family Raphistomatidae.
Cerithiella acutaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Cerithiella.
Cerithiscala parvibrazosaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Cerithiscala tehuacanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Chemnitzia silvaiSp. novValidLandau & MicaliPlioceneA species of Chemnitzia.
Chevallieria texanopsisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Chicomurex parvus[103] Sp. novValidMerle, Landau & BreitenbergerMioceneA species of Chicomurex.
Chicoreus javanusSp. novValidMerle, Landau & BreitenbergerMioceneA species of Chicoreus.
Chicoreus martiniSp. novValidMerle, Landau & BreitenbergerMioceneA species of Chicoreus.
Chicoreus solidusSp. novValidMerle, Landau & BreitenbergerMioceneA species of Chicoreus.
Clava (Semivertagus) scribaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Clavatula hirmetzli[104] Sp. novValidKovács & ViciánMiddle MioceneLajta Limestone FormationA species of Clavatula.
Clavatula santhaiSp. novValidKovács & ViciánMiddle MioceneLajta Limestone FormationA species of Clavatula.
Clavatula szekelyhidiaeSp. novValidKovács & ViciánMiddle MioceneLajta Limestone FormationA species of Clavatula.
Cochlespira (Tahusyrinx) laevisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Cochlespira.
Conilithes herodus[105] Sp. novValidPsarras, Koskeridou & MerleMiocene (Tortonian)A species of Conilithes.
Conradconfusus nodulinus[106] Sp. novValidGarvieEoceneCook Mountain Formation
Conradconfusus texanusSp. novValidGarviePaleocene (Danian)Kincaid Formation
Conus (Kalloconus) asterousiaensisSp. novValidPsarras, Koskeridou & MerleMiocene (Tortonian)
A species of Conus.
Conus (Kalloconus) helladicusSp. novValidPsarras, Koskeridou & MerleMiocene (Tortonian)A species of Conus.
Cordieria biouesensis[107] Sp. novValidPacaudPaleocene (Thanetian)A species of Cordieria.
Coronia argillaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Costoanachis venzoi[108] Nom. novValidHarzhauser & LandauMioceneA species of Costoanachis; a replacement name for Anachis (Costoanachis) subcorrugata Venzo & Pelosio (1963).
CrassaseilaGen. et 2 sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species C. ripleyana and C. paleocenica.
Crassispira plioibericostricta[109] Sp. novValidBrunetti & ForliPliocene (Zanclean)A species of Crassispira.
Crassispira pseudoglabraSp. novValidBrunetti & ForliPliocene (Piacenzian)A species of Crassispira.
Crenaturricula minutaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Creonella hebetertiaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Cretatortulosa cretakachinensisSp. novIn pressYu, Salvador & JarzembowskiLate Cretaceous (Cenomanian)Burmese amber
Cretatortulosa gignens[110] [111] [112] Sp. novIn pressJochum, Yu & NeubauerLate Cretaceous (Cenomanian)Burmese amberA member of the family Pupinidae.
Cryptochorda paleocaenicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Cyathopoma pingyiensis[113] Sp. novIn pressYu et al.CretaceousPaleogene transitionPingyi BasinA species of Cyathopoma.
Cylichna (Cylichnopsis) ellipticaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Cylichna.
Cymatiella dulaiiSp. novValidVicián & KovácsEocene (Lutetian)Csernye FormationA species of Cymatiella.
Cymatiella tzankoviNom. novValidVicián & KovácsPaleoceneA species of Cymatiella; a replacement name for Eutritonium (Sassia) rutoti Tzankov (1940).
Cymatium danianumSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Cymatium.
Cyphoma guraboensis[114] Sp. novValidGroves & LandauEarly PlioceneGurabo FormationA species of Cyphoma.
Cypraeorbis alisonkayaeSp. novValidGroves & LandauMioceneA species of Cypraeorbis.
CyprioconchaGen. et sp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A member of the family Hokkaidoconchidae. The type species is C. robertsoni.
DefensinaGen. et comb. novValidHarzhauser & LandauMiocene

A member of the family Columbellidae. The type species is "Columbella (Mitrella)" bucciniformis Hoernes & Auinger (1880).
Dermomurex wanneriNom. novValidMerle, Landau & BreitenbergerMioceneA species of Dermomurex; a replacement name for Murex acuticostatus Wanner & Hahn (1935).
Desbruyeresia kambiaensisSp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A species of Desbruyeresia.
Desbruyeresia kinousaensisSp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A species of Desbruyeresia.
Desbruyeresia memiensisSp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A species of Desbruyeresia.
Discotectonica paracutissimaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Discotectonica.
Doliella parnitensSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Doliella.
Domiporta amoenaSp. novValidHarzhauser & LandauMioceneDej Formation

A species of Domiporta.
Domiporta pulchraSp. novValidHarzhauser & LandauMioceneDej FormationA species of Domiporta.
Domiporta turpisSp. novValidHarzhauser & LandauMioceneBaden FormationA species of Domiporta.
EderazygaGen. et sp. et comb. novValidPieroni, Monari & ToddLate Triassic (Carnian to Rhaetian)Nayband Formation
San Cassiano Formation
Zu Limestone


A possible Zygopleuridae member.
The type species is E. fanchini;
genus also includes "Cerithium"? lateplicatum Klipstein (1843).
Ellobium mizutanii[115] Sp. novValidKawase & IchiharaEarly MioceneAkeyo FormationA species of Ellobium.
Entemnotrochus kathiawarensis[116] Sp. novValidBose, Das & MondalMiocene (Burdigalian)A species of Entemnotrochus.
Eoacteon paracutissimaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Eoancilla lapicidinaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Eocithara (?) laurenceiSp. novValidGarviePaleocene (Danian)Kincaid Formation
Eopleurotoma atacicaNom. novValidPacaudEocene (Ypresian)A species of Eopleurotoma; a replacement name for Pleurotoma (Eopleurotoma) romani Doncieux (1908).
Eotrivia pacaudi[117] Sp. novValidFehseEocene
Eoturris fallsensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Eoworthenia frydaiNom. novValidKarapunar & NützelA member of Pleurotomariida; a replacement name for Worthenia rarissima Barrande.
Episcomitra antibellardiiSp. novValidHarzhauser & LandauMioceneDej FormationA species of Episcomitra.
Episcomitra leopoldianaSp. novValidHarzhauser & LandauMioceneBaden Formation

A species of Episcomitra.
Episcomitra missileSp. novValidHarzhauser & LandauMioceneGrund Formation
A species of Episcomitra.
Episcomitra neubaueriSp. novValidHarzhauser & LandauMioceneDej FormationA species of Episcomitra.
Episcomitra pontonxensisSp. novValidLozouetOligocene (Chattian)A species of Episcomitra.
Episcomitra pseudoincognitaSp. novValidHarzhauser & LandauMioceneDej Formation
A species of Episcomitra.
Episcomitra vermeuleni[118] Sp. novValidPacaudEocene (Priabonian)A species of Episcomitra.
Eulimella ariejansseniSp. novValidLandau & MicaliPlioceneA species of Eulimella.
Euthema myanmarica[119] Sp. novValidBalashovCretaceousHkamti amberA species of Euthema.
Euthria mellianensisSp. novValidLozouetMioceneA species of Euthria.
Euthria tarusatensisSp. novValidLozouetOligocene (Chattian)A species of Euthria.
Euthriofusus argillensisSp. novValidLozouetOligocene (Chattian)
Falsifusus problematicusSp. novValidGarviePaleocene (Danian)Kincaid Formation
FedosoviaGen. et comb. novValidHarzhauser & LandauMiocene

A member of the family Costellariidae. The type species is "Mitra (Callithea)" fuchsi Hoernes & Auinger (1880).
FraudizibaGen. et nom. et sp. novValidHarzhauser & LandauMiocene and PlioceneKaraman Basin








A member of the family Mitridae. The type species is F. paratethyca (a replacement name for Mitra austriaca Csepreghy-Meznerics, 1950); genus also includes new species F. ottomanica.
Fulgurofusus spinosusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Fulgurofusus.
Fusulculus texanusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Fusulculus.
Galba prima[120] Sp. novValidYu, Neubauer & JochumCretaceousBurmese amberA species of Galba.
Gemmula bearrizensisNom. novValidPacaudEocene (Priabonian)A species of Gemmula; a replacement name for Drillia pellati Boussac (1911).
Gemmula oscaSp. novValidPacaudEocene (Bartonian)A species of Gemmula.
GentilifulgurGen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species G. praemortoni.
Gibbula lovellreeveiNom. novValidHarzhauserPlioceneA Trochidae member;
a replacement name for Trochus (Gibbula) reevei Harmer (1923).
Gibbula pizzolatoi[121] Sp. novValidSpadiniPlioceneA species of Gibbula.
Gibbula steiningeriNom. novValidHarzhauserEarly MioceneA Trochidae member;
a replacement name for Trochus amedei bicincta Schaffer (1912).
Gibbula tavaniiNom. novValidHarzhauserMioceneA Trochidae member;
a replacement name for Gibbula minima Tavani (1939).
Gibbuliculus[122] Gen. et comb. et nom. novValidHarzhauserOligocene to PleistoceneA Trochidae member;
The type species is "Gibbula" brebioni Landau, Van Dingenen & Ceulemans (2017).
Genus includes G. saccoi (a replacement name for Gibbula protumida Sacco, 1896).
"Gibbulinella" sandbergeriNom. novValidHarzhauser & NeubauerEoceneA member of the family Streptaxidae; a replacement name for Pupa simplex Sandberger (1870).
Haedropleura dellabellaiSp. novValidBrunetti & CrestiPlioceneA species of Haedropleura.
Hammatospira cancellata[123] Sp. novValidMazaevEarly Carboniferous
Hespererato canaeSp. novValidGroves & LandauLate Miocene and early PlioceneCercado FormationA species of Hespererato.
Hespererato praeclusaSp. novValidGroves & LandauEarly–middle MioceneBaitoa FormationA species of Hespererato.
Hexaplex nasciumSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Hexaplex.
Hirsuticyclus canaliculatus[124] Sp. novIn pressYuCretaceousBurmese amberA member of the family Cyclophoridae.
Hochheimia[125] Nom. novValidHarzhauser & NeubauerOligoceneA member of the family Hygromiidae; a replacement name for Palaeotrichia Nordsieck (2014).
Hokkaidoconcha morisseauiSp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A member of the family Hokkaidoconchidae.
Holospira? fallsensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Possibly a species of Holospira.
?Hydrocena praecursor[126] Sp. novIn pressYu & NeubauerLate Cretaceous (Cenomanian)Burmese amberA member of the family Hydrocenidae.
Jponsia lacetania[127] Sp. novValidPacaud & GoretEocene (Ypresian)A member of the family Pachychilidae.
Kapalmerella gelusSp. novValidGarviePaleocene (Danian)Kincaid Formation
Keepingia elongataSp. novValidLozouetOligocene (Chattian)
LaevispiraGen. et comb. et sp. novValidRavenMiocene and Pliocene

A member of the family Strombidae. Genus includes "Dolomena" bruneiensis Harzhauser, Raven & Landau (2018), "Strombus" glaber Martin (1879), "Strombus" karikalensis Cossmann (1903), "Strombus" triangulatus Martin (1879) and "Strombus" varinginensis Martin (1899), as well as new species L. seriaensis.
Laevitrivia[128] Gen. et comb. et 5 sp. novValidFehse & ViciánMiocene and Pliocene
A member of the family Triviidae. The type species is "Cypraea" pisolina Lamarck, (1810); genus also includes new species L. annae, L. bertaccinii, L. pliopisolina, L. pseudocalva and L. ventricostata.
Latirus? asperpellisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Possibly a species of Latirus.
Laubella subsulcataSp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida.
LineacingulumGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" texturata Münster (1841).
LineaetomariaGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" decorata Münster (1841).
Liostomia wilvanderstoelae[129] Sp. novValidLandau & MicaliPlioceneA species of Liostomia.
Littorina elongataSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Littorina.
Littorina (Prosthenodon) incertusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Littorina.
Lupira danianumSp. novValidGarviePaleocene (Danian)Kincaid Formation
Lutema gracilisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Lyria elongataSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Lyria.
Lyria tehuacanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Lyria.
Malea hyaduckiSp. novValidPetuch & BerschauerPliocene
A species of Malea.
MartaiaGen. et comb. novValidHarzhauser & LandauMiocene




A member of the family Columbellidae. The type species is "Columbella (Anachis)" dujardini Hoernes & Auinger (1880); genus also includes "Columbella (Anachis)" zitteli Hoernes & Auinger (1880).
Mathilda (Echinimathilda) duogentaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Mathilda.
Melongena pontonxensisSp. novValidLozouetOligocene (Chattian)A species of Melongena.
Metula bulbiformisSp. novValidLozouetOligocene (Chattian)A species of Metula.
Metula nanaSp. novValidLozouetOligocene (Chattian)A species of Metula.
Metula pseudomajorSp. novValidLozouetOligocene (Chattian)A species of Metula.
Metula rivulisensisSp. novValidLozouetOligocene (Chattian)A species of Metula.
Mimospira llangynogensisSp. novValidEbbestad & CopeOrdovician (Floian)A member of the family Clisospiridae.
Mitrella dacicaSp. novValidHarzhauser & LandauMioceneDej Formation

A species of Mitrella.
Mitrella demaintenonaeSp. novValidHarzhauser & LandauMioceneBaden Formation
Dej Formation

A species of Mitrella.
Mitrella elongatissimaSp. novValidHarzhauser & LandauMioceneBaden Formation
A species of Mitrella.
Mitrella fortisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Mitrella.
Mitrella tarbellianaSp. novValidLozouetOligocene (Chattian)A species of Mitrella.
Mitrella viennensisSp. novValidHarzhauser & LandauMioceneBaden Formation




A species of Mitrella.
Moniquia astibiaiSp. novValidPacaud & GoretEocene (Priabonian)A member of the family Pachychilidae.
Monoplex s.l. szakonyiiSp. novValidVicián & KovácsEocene (Lutetian)Csernye FormationA member of the family Cymatiidae.
Mulderia mulderiSp. novValidLandau & MicaliPlioceneA member of the family Pyramidellidae.
MultitrochiaGen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species M. tehuacana.
Nannamoria alquezae[130] Sp. novValidHawkeMioceneA species of Nannamoria.
Nannamoria cadellaSp. novValidHawkeMioceneCadel FormationA species of Nannamoria.
Nannamoria costatumSp. novValidHawkePlioceneCameron Inlet FormationA species of Nannamoria.
Nannamoria flindersiSp. novValidHawkePlioceneCameron Inlet FormationA species of Nannamoria.
Nannamoria gnotukiSp. novValidHawkeA species of Nannamoria.
Nannamoria hiscockiSp. novValidHawkePlioceneCameron Inlet FormationA species of Nannamoria.
Nannamoria maloneiSp. novValidHawkeA species of Nannamoria.
Nannamoria persimilisSp. novValidHawkeMioceneA species of Nannamoria.
Naria praehelvola[131] Sp. novValidFehse & ViciánMioceneA species of Naria.
Natica (Carinacca) kincaidensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Natica.
Nebularia soliphilaSp. novValidHarzhauser & LandauMioceneBaden FormationA species of Nebularia.
Nemaspira[132] Gen. et comb. et 2 sp. novValidMazaevEarly Carboniferous to Late Triassic




?
A member of the family Gosseletinidae. Genus includes several species formerly assigned to the genera Gosseletina and Globodoma, as well as new species N. shamaevi and N. skuini.
Nitidiclavus senesi[133] Sp. novValidBiskupičMioceneA member of the family Drilliidae.
Niveria callosa[134] Sp. novValidFehseLate OligoceneA species of Niveria.
Niveria novosperaraSp. novValidFehseLate OligoceneA species of Niveria.
NodocingulumGen. et comb. et 2 sp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida. The type species is "Pleurotomaria" coronata Münster (1841); genus also includes new species N. ernstkittli and possibly also N? turris.
Nodolargena[135] Gen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
A member of the new family Nodolargenidae. Genus includes new species N. frosti.
NodovolutaGen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species N. coronata.
Odostomia malagensisSp. novValidLandau & MicaliPlioceneA species of Odostomia.
OligonaliaGen. et comb. novValidLozouetOligoceneA new genus for "Siphonalia" riparia Lozouet (1999).
OlinderiaGen. et comb. et 3 sp. novValidLozouetOligocene (Chattian)A member of the family Charitodoronidae. The type species is "Charitodoron" tauzini Lozouet (1991); genus also includes new species O. lestrillensis, O. poustagnacensis and O. verdunensis.
Olivella tehuacanensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Olivella.
Ondina pinguisSp. novValidLandau & MicaliPlioceneA species of Ondina.
Onustus undosusSp. novValidRavenMioceneA species of Onustus.
Opalia tehuacanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Opalia.
Orbitestella kieliSp. novValidChernyshev & GoedertLate Eocene/early OligoceneGries Ranch Formation
A member of the family Orbitestellidae.
Orthosurcula ethaniSp. novValidGarvieEoceneCook Mountain Formation
Oxyacrum gracilisSp. novValidGarviePaleocene (Danian)Kincaid Formation
PagoniaGen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species P. spiralis.
Palaeorhaphis? laevisSp. novValidGarviePaleocene (Danian)Kincaid Formation
PaleofususGen. et 3 sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species P. bellus, P. parvicrassus and P. elegantissima.
Papuliscala? acusSp. novValidGarviePaleocene (Danian)Kincaid Formation
Possibly a species of Papuliscala.
Papuliscala keaniSp. novValidGarvieEoceneCook Mountain Formation
Parasassia vargaiSp. novValidVicián & KovácsEocene (Lutetian)Csernye FormationA member of the family Cymatiidae.
Parthenina pacaudiNom. novValidLandau & MicaliMioceneA species of Parthenina; a replacement name for Pyrgulina (Parthenina) dollfusi Cossmann (1921).
Parvisipho? spiralisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Paskentana dixoniSp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A member of Abyssochrysoidea belonging to the family Paskentanidae.
Paskentana xenophontosiSp. novValidKaim et al.Late Cretaceous (Cenomanian to Turonian)A member of Abyssochrysoidea belonging to the family Paskentanidae.
Perotrochus bermotiensisSp. novValidBose, Das & MondalOligoceneA species of Perotrochus.
Perrona harzhauseriSp. novValidKovács & ViciánMiddle MioceneLajta Limestone FormationA species of Perrona.
Perrona nemethiSp. novValidKovács & ViciánMiddle MioceneLajta Limestone FormationA species of Perrona.
Personopsis merleiSp. novValidVicián & KovácsEocene (Lutetian)Csernye FormationA species of Personopsis.
PfefferiolaNom. novValidHarzhauser & NeubauerEoceneA member of the family Streptaxidae; a replacement name for Oppenheimiella Pfeffer (1930).
Phos estotiensisSp. novValidLozouetOligocene (Chattian)A species of Phos.
Pleurotomella irminonvillaNom. novValidPacaudEocene (Bartonian)A species of Pleurotomella; a replacement name for Pleurotomella cossmanni Morellet & Morellet (1946).
Pliciscala obliquaSp. novValidGarvieEoceneCook Mountain Formation
Poirieria (Paziella) deleraSp. novValidGarviePaleocene (Danian)Kincaid Formation
Polinices tehuacanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Polinices.
Popenoeum tabulatumSp. novValidGarviePaleocene (Danian)Kincaid Formation
PressulasphaeraGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" pamphilus d'Orbigny (1850).
Priscoficus johnfoxiSp. novValidGarviePaleocene (Danian)Kincaid Formation
Priscoficus nodulinaSp. novValidGarviePaleocene (Danian)Kincaid Formation
ProceraedisGen. et 2 sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species P. imbricataria and P. tegumenis.
Prososthenia esuegirottii[136] Sp. novValidDominici & ForliPliocene (Zanclean)A member of the family Hydrobiidae.
Protoplex? zsoldosiSp. novValidVicián & KovácsEocene (Lutetian)Csernye FormationA member of the family Cymatiidae.
Proviviparus[137] Gen. et sp. novValidFrese & PonderLate JurassicTalbragar Fish BedA member of the family Viviparidae. Genus includes new species P. talbragarensis
PseudoananiasGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" subgranulata Münster (1841).
Pseudokatosira uhlhingleyorum[138] Sp. novValidGründel, Davies & SimonsenEarly Jurassic (Pliensbachian)
Pseudolatirus acuelatusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Pseudolatirus.
Pseudolatirus boettgeri[139] Nom. novValidKovácsMioceneA member of the family Fasciolariidae; a replacement name for Fusus sublamellosus Boettger (1906). Originally described as a species of Pseudolatirus; Harzhauser, Landau & Vermeij (2024) tentatively placed it in the genus Takashius.[140]
PseudosassiaGen. et comb. et 2 sp. novValidVicián & KovácsPaleocene–Miocene















A member of the family Cymatiidae. The type species is "Triton" flandricum Koninck (1838); genus also includes new species P. gurdoni and P. traceyi.
Pseudoterebellum[141] Gen. et comb. novValidMaxwell, Rymer & CongdonEoceneGhazij Shales

A member of the family Seraphsidae. The type species is "Terebellum" diversiornatum Eames (1952); genus also includes "Terebellum" saudiensis Abbass (1972) and "Terebellum" olympiae Rolando (2001).
Ptychatractus cataractusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Ptychatractus.
Pulchraspira[142] Gen. et sp. novIn pressYu, Salvador & JarzembowskiLate Cretaceous (Cenomanian)Burmese amberGenus includes new species P. teneristoma.
Puposyrnola? confusaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Possibly a species of Puposyrnola.
Puposyrnola multibacaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Puposyrnola.
Puposyrnola obliquaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Puposyrnola.
Puposyrnola shirleyaeSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Puposyrnola.
Pusia aturensisSp. novValidLozouetOligocene (Chattian)A species of Pusia.
Pusia confundaSp. novValidHarzhauser & LandauMioceneDej FormationA species of Pusia.
Pusia? falsitranssylvanicaNom. novValidHarzhauser & LandauMioceneDej FormationPossibly a species of Pusia; a replacement name for Mitra (Costellaria) recticosta var. transsylvanica Boettger (1902).
Pusia subpyrenaicaSp. novValidLozouetOligocene (Chattian)A species of Pusia.
Pusia virodunensisSp. novValidLozouetOligocene (Chattian)A species of Pusia.
Pustularia korneliSp. novValidFehse & ViciánMioceneA species of Pustularia.
Pyrgiscus elegans[143] Sp. novValidTabanelli et al.PlioceneA species of Pyrgiscus.
Pyrgiscus jaapmulderiSp. novValidLandau & MicaliPlioceneA species of Pyrgiscus.
Pyrgulina marliesaeSp. novValidLandau & MicaliPlioceneA species of Pyrgulina.
Retusa (Cylichnina) praemoryiSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Retusa.
Rhaphaulus zhuoiSp. novIn pressYu, Salvador & JarzembowskiLate Cretaceous (Cenomanian)Burmese amber
Rinaldoella tornataSp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida.
Ringicula taeniaSp. novValidGarvieEoceneCook Mountain Formation
A species of Ringicula.
Sansonia texanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
SassellaGen. et 2 sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species S. mixta and S. frosti.
Scalptia nemethi[144] Sp. novValidKovács & ViciánMioceneLajta FormationA species of Scalptia.
Scaphella kendrae[145] Sp. novValidPetuch & BerschauerPliocene (Zanclean)Murdock Formation
A species of Scaphella.
Schizogonium undaeSp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida.
Seraphs kaindraperiSp. novValidMaxwell, Rymer & CongdonEocene (Priabonian)A member of the family Seraphsidae.
SimilalvaniaGen. et comb. novValidChirli & ForliPlioceneA member of the family Rissoidae. The type species is "Alvania" magistra Chirli (2006)
Solariella microstriataSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Solariella.
Solariorbis planulatisSp. novValidGarviePaleocene (Danian)Kincaid Formation
SquamowortheniaGen. et comb. novValidMazaevCarboniferous

Genus includes S. duponti (Holzapfel, 1889) and "Worthenia" crenilunula Yoo (1994).
Stellaria striatissimaSp. novValidRavenMioceneA species of Stellaria.
StriacingulumGen. et comb. novValidKarapunar & NützelA member of Pleurotomariida. The type species is "Pleurotomaria" cancellatocingulata Klipstein (1844).
Striaticostatum figusSp. novValidGarviePaleocene (Danian)Kincaid Formation
Stuckenbergispira dombarensisSp. novValidMazaevEarly Carboniferous
Sveltella novusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Sveltella.
Sveltia sofiae[146] Sp. novValidLandau & da SilvaPlioceneA species of Sveltia.
Syrnola debraeSp. novValidGarvieEoceneCook Mountain Formation
A species of Syrnola.
Taioma? argillaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Taioma calxaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Taranis occultaSp. novValidBrunetti & CrestiPlioceneA species of Taranis.
Teinostoma regularisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Teinostoma.
Terebellum humilispirumSp. novValidRavenMioceneA species of Terebellum.
Terebra (Mirula) mexiaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Terebra.
Terebrifusus megahaniSp. novValidGarvieEoceneCook Mountain Formation
Thylacodes devriesi[147] Sp. novIn pressSanfilippo et al.MioceneA species of Thylacodes.
Tornatellaea globosaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Tornatellaea liniferaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Tripsycha strappleiSp. novValidGarviePaleocene (Danian)Kincaid Formation
Tritia miniridibundaSp. novValidLozouetMioceneA species of Tritia.
Tritia praebasterotiSp. novValidLozouetOligocene (Chattian)A species of Tritia.
Tritiaria formosaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Trivia disjunctaSp. novValidFehseLate OligoceneA species of Trivia.
Trivia simplexSp. novValidFehseLate OligoceneA species of Trivia.
Trona colossusSp. novValidGroves & LandauEarly–middle MioceneBaitoa FormationA species of Trona.
Truncatella jiaozhouensis[148] Sp. novIn pressYu et al.Late CretaceousJiaolai BasinA species of Truncatella.
TumidosiphoGen. et sp. novValidGarviePaleocene (Danian)Kincaid Formation
Genus includes new species T. tehuacana.
Turbo (Marmarostoma) ishidai[149] Sp. novValidTomida, Sano & KaseMioceneEna LimestoneA species of Turbo.
Turbonilla binckiSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turbonilla bongiardinoiSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turbonilla crovatoiSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turbonilla malacitanaSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turbonilla mauroiSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turbonilla plioalboranensisSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turbonilla velerinensisSp. novValidLandau & MicaliPlioceneA species of Turbonilla.
Turricula frostiSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Turricula.
Turricula imbricataSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Turricula.
"Turritella" calaxSp. novValidGarviePaleocene (Danian)Kincaid Formation
Umbilia furneauxensis[150] Sp. novValidSouthgate, Militz & RobertsPlioceneA species of Umbilia.
Unitas duomilliasSp. novValidGarvieEoceneCook Mountain Formation
Unitas euaniSp. novValidGarviePaleocene (Danian)Kincaid Formation
Unitas petitiSp. novValidGarviePaleocene (Danian)Kincaid Formation
Valvata jiaolaiensisSp. novIn pressYu et al.Late CretaceousJiaolai BasinA species of Valvata.
Vanikoropsis bashforthi[151] Sp. novValidSchnetler & NielsenPaleocene (Danian)Agatdal FormationPossibly a member of the family Vanikoridae.
Vanikoropsis jakobseniSp. novValidSchnetler & NielsenPaleocene (Danian)Agatdal FormationPossibly a member of the family Vanikoridae.
Vanikoropsis mortenseniSp. novValidSchnetler & NielsenPaleocene (Selandian)Kerteminde MarlPossibly a member of the family Vanikoridae.
Varicospira bekenuensisSp. novValidRavenMioceneA species of Varicospira.
Varicospira reticulataSp. novValidRavenMioceneA species of Varicospira.
Vermicularia? columellaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Vexillum transalpinumSp. novValidHarzhauser & LandauMioceneBaden Formation


A species of Vexillum.
Vitrinella discusSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Vitrinella.
Vitrinella praelaevisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Vitrinella.
Volutifusus dougsheltoniSp. novValidPetuch & BerschauerPliocene (Piacenzian)Tamiami Formation
A species of Volutifusus.
Volutifusus kissimmeensisSp. novValidPetuch & BerschauerPliocene (Piacenzian)Tamiami Formation
A species of Volutifusus.
Volutomorpha ultimaSp. novValidGarviePaleocene (Danian)Kincaid Formation
WormsinaGen. et comb. novValidHarzhauser & LandauMioceneA member of the family Mitridae. The type species is "Mitra (Cylindra)" transsylvanica Hoernes & Auinger (1880).
Wortheniella klipsteiniSp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida.
Wortheniella paolofedeleiSp. novValidKarapunar & NützelLate TriassicSan Cassiano FormationA member of Pleurotomariida.
Zafrona sphaerocorrugataSp. novValidHarzhauser & LandauMioceneBaden Formation




A species of Zafrona.
Zebinella tehuacanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Zebinella.
Zonaria vokesaeSp. novValidGroves & LandauEarly–middle MioceneBaitoa FormationA species of Zonaria.

Bivalves

New taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages
Acar beadataSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Acar.
Amphidonte mirkamalovi[152] Sp. novValidMetelkin & KosenkoEarly Cretaceous (Aptian)An oyster.
Arcopsis subrhombusSp. novValidGarviePaleocene (Danian)Kincaid Formation
Arcopsis tumulusSp. novValidGarviePaleocene (Danian)Kincaid Formation
Arkhanguelskya[153] Gen. et comb. novValidCooper & LeanzaCretaceousA member of the family Buchotrigoniidae. Genus includes "Trigonia" weberi Arkhangelsky (1916).
Baluchicardia? eoplataSp. novValidGarviePaleocene (Danian)Kincaid Formation
Bengtsonigonia[154] Nom. novValidCeccolini & CianferoniEarly CretaceousRiachuelo FormationA replacement name for Bengtsonella Cooper & Leanza (2019).
BohemigoniaGen. et sp. et comb. novValidCooperA member of the family Megatrigoniidae. The type species is B. frici; genus also includes "Trigonia" debuchii Geinitz (1840).
Caryocorbula? praefigeraSp. novValidGarviePaleocene (Danian)Kincaid Formation
Chama ferrata[155] Sp. novValidBerezovskyEoceneA species of Chama.
Chama subcalcarataSp. novValidBerezovskyEoceneA species of Chama.
Corbulamella palaeocenicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Crassatellina minuta[156] Sp. novValidJaitly et al.Late Cretaceous (Turonian)Nodular Limestone FormationA member of the family Lucinidae belonging to the subfamily Eriphylinae.
Crassatellina paleocaenicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Cuspidaria texanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Cuspidaria.
Diplodonta frostiSp. novValidGarviePaleocene (Danian)Kincaid Formation
Diplodonta paleocaenicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
DorbigoniaGen. et comb. novValidCooperCretaceousA member of the family Megatrigoniidae. Genus includes "Trigonia" disparilis d'Orbigny (1844).
ElianegoniaNom. novValidCeccolini & CianferoniLate CretaceousA replacement name for Elianella Cooper (2015).
Eomiltha strangensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Freneixicardia picturata[157] Sp. novValidBerezovskyEoceneA cockle.
Frenguelliella eopacifica[158] Sp. novValidEchevarría, Damborenea & ManceñidoEarly JurassicA member of Trigoniida belonging to the family Frenguelliellidae.
Glycymeris laevigataSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Glycymeris.
Grammatodon superstesSp. novValidGarviePaleocene (Danian)Kincaid Formation
Gregariella frostiSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Gregariella.
Himenourella[159] Gen. et comb. novValidCooperCretaceous
A member of the family Megatrigoniidae. Genus includes "Trigonia" jimbonis Strand (1928), "Trigonia" sawatai Yehara (1923), "Apiotrigonia" subjimboi Miroljubov in Vereshchagin et al. (1965) and "Trigonia" subovalis Jimbo (1894).
Kolymia rutskovi[160] Sp. novValidBiakov in Biakov et al.Permian
KushmurunskyaGen. et comb. novValidCooperCretaceousA member of the family Megatrigoniidae. Genus includes "Trigonia" gladkowskii Shcheglova-Borodina (1960).
KyzylkumgoniaGen. et comb. novValidCooperCretaceousA member of the family Megatrigoniidae. Genus includes "Trigonia" syrdariensis Arkhangelsky (1916), "Trigonia" turcmenensis Arkhangelsky (1916) and "Megatrigonia" tagamensis Belyakova (1980).
Loxocardium marmoreumSp. novValidBerezovskyEoceneA cockle.
Lucina crassiplicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Lucina.
Lucina multivittaeSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Lucina.
Lucina tehuacanensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Lucina.
MaedagoniaGen. et comb. et nom. novValidCooperCretaceous
A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" mikasaensis Tashiro (1979), "Apiotrigonia" ashizawaensis Saito (1962), "Apiotrigonia" crassoradiata Nakano (1957), "Apiotrigonia" futabaensis Maeda & Kawabe (1967), "Apiotrigonia" hetonaiana Tashiro (1978), "Apiotrigonia" hironoensis Maeda & Kawabe (1967), "Apiotrigonia" jimboi Nakano (1957), "Apiotrigonia" obliquecostata Nakano (1957), "Apiotrigonia" obsoleta Nakano (1957), "Apiotrigonia" orikiensis Maeda & Kawabe (1967), "Apiotrigonia" undulosa Nakano (1957) and "Apiotrigonia" utoensis Tashiro (1972), as well as M. yabei (a replacement name for Trigonia subovalis var. minor Yabe & Nagao, 1925).
Martesia mazanticus[161] Sp. novValidCastañeda-Posadas, Trujillo-Hernández & Zúñiga-MijangosMioceneMexican amberA species of Martesia.
MatsudagoniaGen. et comb. novValidCooperCretaceousA member of the family Megatrigoniidae. Genus includes "Apiotrigonia" dubia Tashiro (1979).
MoerickellaGen. novValidEchevarría, Damborenea & ManceñidoEarly JurassicA member of Trigoniida, most likely the oldest member of the family Myophorellidae.
Naiadites devonicus[162] Sp. novValidScholze & GessDevonian (Famennian)Witpoort Formation
Opis reticulataSp. novValidJaitly et al.Late Cretaceous (Turonian)Nodular Limestone FormationA member of the family Lucinidae belonging to the subfamily Opinae.
Ostrea jibananandai[163] Sp. novValidHalder & MitraEocene (Ypresian)Cambay ShaleA species of Ostrea.
Palaeolignopholas[164] Gen. et sp. novValidBolotov et al.Late Cretaceous (Cenomanian)Burmese amberA member of the family Pholadidae belonging to the subfamily Martesiinae. The type species is P. kachinensis.
Pandora? paleocenicaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Parmicorbula? sublaevisSp. novValidGarviePaleocene (Danian)Kincaid Formation
PerezigoniaNom. novValidCeccolini & CianferoniMiddle JurassicA replacement name for Perezella Cooper (2015).
Pojetaia robsonae[165] Sp. novValidPeel & SkovstedCambrian Stage 4Kane Basin Formation
Porterius? strangensisSp. novValidGarviePaleocene (Danian)Kincaid Formation
PoultoniellaGen. et comb. novValidCooperCretaceous
A member of the family Megatrigoniidae. Genus includes "Trigonia" newcombei Packard (1921).
Promyophorella basoaltorumSp. novValidEchevarría, Damborenea & ManceñidoEarly JurassicA member of Trigoniida belonging to the family Myophorellidae.
Promyophorella? sanjuaninaSp. novValidEchevarría, Damborenea & ManceñidoEarly JurassicA member of Trigoniida belonging to the family Myophorellidae.
PseudapiotrigoniaGen. et comb. novValidCooper
(
)
A member of the family Vaugoniidae. Genus includes "Trigonia" sturgisensis Whitfield & Hovey (1906), as well as "Vaugonia" utahensis Imlay (1964).
PseudovaugoniaGen. novValidEchevarría, Damborenea & ManceñidoEarly JurassicA member of Trigoniida belonging to the family Myophorellidae.
Pteromeris nodulinaSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Pteromeris.
RepmanigoniaNom. novValidCeccolini & CianferoniJurassicA replacement name for Repmania Kolesnikov (1980).
Schedocardia imperfectaSp. novValidBerezovskyEoceneA cockle.
SinoliaoningiaNom. novValidCeccolini & CianferoniLate TriassicYangcaogou FormationA replacement name for Liaoningia Yu & Dong (1993).
SokolovskyaGen. et comb. novValidCooperA member of the family Megatrigoniidae. Genus includes "Apiotrigonia" levinae Pojarkova (1990) and "Apiotrigonia" khoresmensis Belyakova (1965).
SonoragoniaGen. et comb. et nom. novValidCooperEarly CretaceousLowell Formation
A member of the family Megatrigoniidae. Genus includes "Trigonia" cragini Stoyanow (1949), as well as S. scotti (a replacement name for Trigonia kitchini Stoyanow, 1949).
Sphenotomorpha orientalis[166] Sp. novValidDalenz FarjatDevonian (Pragian)Icla FormationA member of Modiomorphida belonging to the family Modiomorphidae.
Sunetta crassatelliformis[167] Sp. novValidHaga & Fukuda in Fukuda et al.Early PleistoceneA species of Sunetta.
Sunetta nomuraiSp. novValidHaga & Fukuda in Fukuda et al.PleistoceneA species of Sunetta.
Syndesmyella pliominor[168] Sp. novValidBrunetti & CrestiEarly PlioceneA member of the family Semelidae.
Tellina brevisSp. novValidGarviePaleocene (Danian)Kincaid Formation
A species of Tellina.
Terrypoultoniella[169] Gen. et sp. novValidEchevarría, Damborenea & ManceñidoEarly Jurassic (PliensbachianToarcian)
?
A member of Trigoniida belonging to the family Frenguelliellidae. The type species is "Poultoniella" jaworskii Echevarría, Damborenea & Manceñido (2021); genus might also include "Trigonia" malladae Choffat (1885).
Tindaria hamuroi[170] Sp. novValidAmano & HagaMioceneHigashibessho FormationA member of the family Tindariidae.
Titomaya[171] Gen. et comb. et sp. novValidAlvarez & del RíoPaleocene (Danian)Lefipán Formation
Salamanca Formation
A member of Venerida belonging to the group Veneroidea. Genus includes "Meretrix" chalcedonica, as well as new species T. longobucca.
Tivelina texanaSp. novValidGarviePaleocene (Danian)Kincaid Formation
Tivelina trigonalisSp. novValidGarviePaleocene (Danian)Kincaid Formation
Unionites kobyumensis[172] Sp. novValidBiakov in Biakov & KutyginLate Permian
Vaccinites alencasteri[173] Sp. novValidMitchellLate Cretaceous (probably late Turonian or early Coniacian)Cuautla FormationA rudist bivalve.
Vaccinites temazcaliSp. novValidMitchellLate Cretaceous (Campanian)Temazcal LimestoneA rudist bivalve.
VillamiliaGen. et comb. novValidCooper & LeanzaCretaceousA member of the family Buchotrigoniidae. Genus includes "Buchotrigonia" etayoi Villamil (1992), "Trigonia" humboldtii Buch (1839), "Buchotrigonia" kauffmani Villamil (1992), "Buchotrigonia" perezi Villamil (1992) and "Vaugonia niranohamensis" santamariae Geyer (1973; raised to the rank of the species V. santamariae).
Weyla santuccii[174] Sp. novValidDamborenea et al.Early Jurassic (Hettangian)Pogibshi Formation
A scallop.
Whitfieldiella luceaeSp. novValidMitchellLate Cretaceous (Campanian)Clifton Limestone
Cotui Limestone


A rudist bivalve.
Yunannia[175] Gen. et 2 sp. novValidZhang & Niu in Zhang et al.OrdovicianDongchong FormationA member of Pteriomorphia of uncertain phylogenetic placement. The type species is Y. gankengensis; genus also includes Y. yunkaiensis.

Research

Other molluscs

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages
Anabarella? navaranae[178] Sp. novValidPeelCambrian Stage 4Aftenstjernesø Formation
Cadulus praepalmeraeSp. novValidGarviePaleocene (Danian)Kincaid Formation
A tusk shell, a species of Cadulus.
Catalanispira primaSp. novValidEbbestad & CopeOrdovician (Floian)A member of the family Onychochilidae.
Figurina? polarisSp. novValidPeelCambrian Stage 4Aftenstjernesø FormationA helcionellid. Subsequently made the type species of a separate genus Sermeqiconus.[179]
Hippopharangites groenlandicusSp. novValidPeelCambrian Stage 4Aftenstjernesø FormationA member of the stem group of Aculifera.
Holoornatus[180] Gen. et sp. novIn pressZhang et al.Cambrian (Fortunian)A maikhanellid. Genus includes new species H. strigatus.
InughuitoconusGen. et sp. novValidPeelCambrian Stage 4Aftenstjernesø FormationA member of the stem group of Aculifera. Genus includes new species I. borealis.
Lepidozona luzanovkensis[181] Sp. novValidSirenkoPaleoceneLuzanovka BedsA chiton, a species of Lepidozona.
Minodentalium[182] Gen. et 2 sp. et comb. novValidAsato & KasePermianAkasaka Limestone


A tusk shell. Genus includes new species M. hayasakai and M. okumurai, as well as "Plagioglypta" furcata Waterhouse (1980), "Pl." girtyi Knight (1940), "Pl." subannulata Easton (1962), "Dentalium" ingens De Koninck (1843), "D." meekianum Geinitz (1866), "Pl." prosseri Morningstar (1922), "Dentalium" priscum Münster in Goldfuss (1842) and "D." herculeum De Koninck (1863).
Ocruranus? kangerlukSp. novValidPeelCambrian Stage 4Aftenstjernesø FormationA member of the stem group of Aculifera.
Prodentalium onoiSp. novValidAsato & KasePermianAkasaka LimestoneA tusk shell.
Progadilinoides[183] Gen. et sp. novValidGuzhovEarly Cretaceous (Aptian)A tusk shell belonging to the family Pulsellidae. The type species is P. balakovensis.
Pseudomyona groenlandica[184] Sp. novValidPeelCambrian (Drumian)Fimbuldal FormationA member of Rostroconchia belonging to the group Tuarangiida and the family Tuarangiidae.
Qingjianglepas[185] Gen. et sp. novIn pressLi et al.CambrianA minute limpet-shaped shell, probably belonging to a helcionelloid. Genus includes new species Q. elegans.
Stenotheca? higginsiSp. novValidPeelCambrian Stage 4Aftenstjernesø Formation

General research

Notes and References

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  2. Korn . D. . Ghaderi . A. . Devaere . L. . Khanehbad . M. . Ammonoids from the Carboniferous-Permian boundary of east-central Iran . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 301 . 3 . 335–345 . 10.1127/njgpa/2021/1015 . 239690567 .
  3. Énay . R. . Mangold . C. . Jurassic ammonites from Central Saudi Arabia (Jebel Tuwaiq and adjacent areas) . 2021 . Revue de Paléobiologie, Genève . 40 . 1 . 1–197 . 10.5281/zenodo.5008788 .
  4. Nakada . K. . Goto . M. . Meister . C. . Matsuoka . A. . The late Pliensbachian (Early Jurassic) ammonoid Amaltheus in Japan: systematics and biostratigraphic and paleobiogeographic significance . 2021 . Journal of Paleontology . 95 . 5 . 1004–1021 . 10.1017/jpa.2021.39 . 2021JPal...95.1004N . 236379379 .
  5. Taylor . D. . Guex . J. . Lucas . S. G. . Ammonoids of the latest Triassic Gabbs Formation at New York Canyon, Mineral County, Nevada . 2021 . New Mexico Museum of Natural History and Science Bulletin . 82 . 393–425 . 2021-01-02 . 2021-02-21 . https://web.archive.org/web/20210221083216/https://www.researchgate.net/publication/348002169_AMMONOIDS_OF_THE_LATEST_TRIASSIC_GABBS_FORMATION_AT_NEW_YORK_CANYON_MINERAL_COUNTY_NEVADA . live .
  6. Taylor . D. . Guex . J. . Early Sinemurian ammonoids and biochronology of the Sunrise Formation, New York Canyon, Mineral County, Nevada . 2021 . New Mexico Museum of Natural History and Science Bulletin . 82 . 371–391 . 2021-01-04 . 2021-02-21 . https://web.archive.org/web/20210221083216/https://www.researchgate.net/publication/348008353_EARLY_SINEMURIAN_AMMONOIDS_AND_BIOCHRONOLOGY_OF_THE_SUNRISE_FORMATION_NEW_YORK_CANYON_MINERAL_COUNTY_NEVADA . live .
  7. Smyshlyaeva . O. P. . Zakharov . Y. D. . New findings of Early Triassic representatives of the suborders Ussuritina and Paraceltitina (Ammonoidea) from south Primorye . 2021 . Paleontological Journal . 55 . 5 . 489–496 . 10.1134/S0031030121050142 . 238640246 .
  8. Söte . T. . Becker . R. T. . Herd . K. J. . Bockwinkel . J. . Upper Frasnian Tornoceratidae (Ammonoidea) from the Sand Formation (Bergisch Gladbach-Paffrath Syncline, Rhenish Massif) . 2021 . PalZ . 95 . 2 . 237–273 . 10.1007/s12542-020-00546-3 . 235220733 . free .
  9. Galácz . A. . Kheraiceras, the Callovian (Middle Jurassic) dimorphic ammonite genus in the South Hungarian Mecsek and Villány Mts. . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 299 . 3 . 275–290 . 10.1127/njgpa/2021/0969 . 233601499 .
  10. Kment . K. . Biostratigraphy of the Hettangian Stage in the Northern Calcareous Alps (Austria, Bavaria) with ammonoids and their classification into biohorizons . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 301 . 3 . 233–282 . 10.1127/njgpa/2021/1007 . 239627293 .
  11. Nannarone . C. . Bilotta . M. . 2021 . Ammoniti titoniani (Zone a Fallauxi e a Volanense) del massiccio Monte Acuto-Monte Catria (Appennino umbromarchigiano): nuovi dati sulla famiglia Simoceratidae Spath, 1924 . Annali del Museo Civico di Rovereto. Sezione: Archeologia, Storia, Scienze Naturali . 37 . 183–234 .
  12. Simms . M. J. . Edmunds . M. . Ammonites from the Lias Group (Lower Jurassic, Sinemurian and Pliensbachian) of White Park Bay, Co. Antrim, Northern Ireland . 2021 . Proceedings of the Geologists' Association . 132 . 6 . 743–763 . 10.1016/j.pgeola.2021.06.001 . 2021PrGA..132..743S . 237768384 .
  13. Wilmsen . M. . Wondrejz . C. . Püttmann . T. . Kennedy . W. J. . Cibolaites petraschecki sp. nov., a new collignoniceratine ammonite from the Brießnitz Formation of Saxony (Turonian, Elbtal Group, Germany) . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 300 . 2 . 189–199 . 10.1127/njgpa/2021/0984 . 236420288 .
  14. Bert . D. . Reboulet . S. . Vernet . B. . Bersac . S. . Canut . L. . Early Crioceratites (heteromorphic ammonites) from the lower Hauterivian of south-eastern France: systematics, intraspecific variation and biostratigraphic implications . 2021 . Cretaceous Research . 126 . Article 104903 . 10.1016/j.cretres.2021.104903 . 2021CrRes.12604903B .
  15. Misaki . A. . Tsujino . Y. . A New Species of the Heteromorph Ammonoid Didymoceras from the Upper Cretaceous Izumi Group in Shikoku, Southwestern Japan, and Its Evolutionary Implications . 2021 . Paleontological Research . 25 . 2 . 127–144 . 10.2517/2020PR010 . 233029551 .
  16. Vermeulen . J. . Leroy . L. . Borro . A. . Lazarin . P. . Mascarelli . E. . 2021 . Nouveau taxa de la famille des Holcodiscidae Spath, 1923 (Ammonitida, Ammonitina, Barremitoidea) . Riviera Scientifique . 105 . 81–96 .
  17. Witts . J. D. . Landman . N. H. . Garb . M. P. . Irizarry . K. M. . Larina . E. . Thibault . N. . Razmjooei . M. J. . Yancey . T. E. . Myers . C. E. . 2021 . Cephalopods from the Cretaceous-Paleogene (K-Pg) boundary interval on the Brazos River, Texas, and extinction of the ammonites . American Museum Novitates . 3964 . 1–52 . 10.1206/3964.1. 0003-0082 . 2246/7250 . 231597207 .
  18. Korn . D. . Revision of Tornoceras typus (Sandberger & Sandberger, 1851) – an iconic Devonian ammonoid of a clade with slow morphological evolution . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 302 . 2 . 147–167 . 10.1127/njgpa/2021/1026 . 243895395 .
  19. Tongtherm . K. . Nabhitabhata . J. . Tuanapaya . S. . Two new Triassic eutomocerid ammonoid cephalopods (Ammonoidea: Ceratitidae) from southeastern peninsular Thailand . 2021 . Phuket Marine Biological Center Research Bulletin . 78 . 147–153 . 10.14456/pmbcrb.2021.11 .
  20. Korn . D. . Bockwinkel . J. . The pharciceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea) . 2021 . European Journal of Taxonomy . 771 . 1–79 . 10.5852/ejt.2021.771.1503 . 239161530 . free .
  21. Zong . P. . Becker . R. T. . Ma . X. . The ammonoid Gonioclymenia fauna from the Upper Devonian (upper Famennian) of western Junggar, northwestern China . 2021 . Geological Journal . 56 . 12 . 5967–5986 . 10.1002/gj.4182 . 236251674 .
  22. Mietto . P. . Jin . X. . Manfrin . S. . Lu . G. . Shi . Z. . Gianolla . P. . Huang . X. . Preto . N. . 2021 . Onset of sedimentation near the Carnian/Norian boundary in the northwestern Sichuan Basin: New evidence from ammonoid biostratigraphy and zircon U–Pb geochronology . Palaeogeography, Palaeoclimatology, Palaeoecology . 567 . Article 110246 . 10.1016/j.palaeo.2021.110246 . 2021PPP...56710246M . 11392/2437547 . 233573583 . free .
  23. Matsukawa . M. . 2021 . Aptian (Lower Cretaceous) ammonite fauna of the Todai Formation, Nagano Prefecture, Japan . Cretaceous Research . 126 . Article 104771 . 10.1016/j.cretres.2021.104771 . 2021CrRes.12604771M .
  24. Polkovoy . K. S. . A new genus Kislovodskoceras (Desmoceratidae, Ammonoidea) from the middle Aptian of the vicinity of Kislovodsk (northern Caucasus) . 2021 . Paleontological Journal . 55 . 5 . 520–528 . 10.1134/S0031030121050129 . 238640207 .
  25. Goy . A. . Comas-Rengifo . M. J. . 2021 . Nejdia (Hildoceratidae, Ammonoidea) in the Toarcian (Lower Jurassic) of the Iberian Peninsula. Systematics, phylogenetic affinities and palaeobiogeography . Historical Biology . 33 . 12 . 3173–3190 . 10.1080/08912963.2020.1853719 . 233815255 .
  26. Jenks . J. F. . Maekawa . T. . Ware . D. . Shigeta . Y. . Brayard . A. . Bylund . K. G. . Late Griesbachian (Early Triassic) ammonoids and nautiloids from the Dinwoody Formation at Crittenden Springs, Elko County, Nevada . 2021 . New Mexico Museum of Natural History and Science Bulletin . 86 . 1–23 .
  27. Gulyaev . D. B. . Ippolitov . A. P. . Lower Callovian of Kanev Dislocations, Cherkasy Oblast, Ukraine: Ammonites and Stratigraphy . 2021 . Stratigraphy and Geological Correlation . 29 . 7 . 767–847 . 10.1134/S0869593821070029 . 2021SGC....29..767G . 245569131 .
  28. Mitta . V. V. . Ammonites of the Ryazanian stage of the Russian Platform: genus Pronjaites Sasonova . 2021 . Paleontological Journal . 55 . 5 . 511–519 . 10.1134/S0031030121050105 . 238640464 .
  29. Jain . S. . Mazur . M. . New dimorphic species of the genus Rollierites Jeannet from southern Poland . 2021 . Annales Societatis Geologorum Poloniae . 91 . 4 . 405–418 . 10.14241/asgp.2021.21 . 246497005 . free .
  30. Samaniego-Pesqueira . A. . Moreno-Bedmar . J. A. . Álvarez-Sánchez . L.E.Ó.N.F. . 2021 . Upper Aptian ammonite biostratigraphy of the Agua Salada and Mural formations, Sonora State, northwest Mexico . Journal of South American Earth Sciences . 112, Part 1 . Article 103558 . 10.1016/j.jsames.2021.103558 . 2021JSAES.11203558S . 244185303 .
  31. Kennedy . W. J. . Latil . J.-L. . Lower Cenomanian ammonites from la Bédoule, Bouches-du-Rhône, France . 2021 . Revue de Paléobiologie, Genève . 40 . 1 . 211–234 . 10.5281/zenodo.5008792 .
  32. Mitta . V. V. . Ammonites and Stratigraphy of the Upper Bajocian Strenoceras niortense Zone in the Interfluve between the Kuban and Urup Rivers (Northern Caucasus) . 2021 . Paleontological Journal . 55 . 12 . 1458–1475 . 10.1134/S0031030121120042 . 247387371 .
  33. Tabrizi . N. G. . Ghaderi . A. . Ashouri . A. R. . Korn . D. . A new record of the Permian ammonoid family Cyclolobidae from Julfa (NW Iran) . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 302 . 2 . 221–230 . 10.1127/njgpa/2021/1029 . 243915631 .
  34. Korn . D. . Revision of Tornoceras frechi Wedekind, 1918 and consequences for the Late Devonian ammonoid stratigraphy . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 300 . 3 . 291–302 . 10.1127/njgpa/2021/0991 . 236231776 .
  35. Lehmann . J. . Bayliss . H. M. . 2021 . The heteromorph ammonite Tropaeum (Tropaeum) evolutum sp. nov. from the Hythe Formation, Lower Greensand Group (basal upper Aptian, Lower Cretaceous) of Maidstone, Kent, UK . Cretaceous Research . 132 . Article 105110 . 10.1016/j.cretres.2021.105110 . 244808527 .
  36. Zaitsev . B. A. . 2021 . Early Jurassic (Late Sinemurian–Early Pliensbachian) Ammonites from the Limestone Boulders of Bodrak River Basin, Southwest Crimea . Stratigraphy and Geological Correlation . 29 . 4 . 411–433 . 10.1134/S0869593821040043 . 2021SGC....29..411Z . 236640510 .
  37. Hoffmann . R. . Slattery . J. S. . Kruta . I. . Linzmeier . B. J. . Lemanis . R. E. . Mironenko . A. . Goolaerts . S. . De Baets . K. . Peterman . D. J. . Klug . C. . 2021 . Recent advances in heteromorph ammonoid palaeobiology . . 96 . 2 . 576–610 . 10.1111/brv.12669 . free . 33438316 .
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  43. Peterman . D. J. . Ritterbush . K. A. . Ciampaglio . C. N. . Johnson . E. H. . Inoue . S. . Mikami . T. . Linn . T. J. . Buoyancy control in ammonoid cephalopods refined by complex internal shell architecture . 2021 . Scientific Reports . 11 . 1 . Article number 8055 . 10.1038/s41598-021-87379-5 . 33850189 . 8044186 . 2021NatSR..11.8055P .
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  45. Hoffmann . R. . Morón-Alfonso . D. . Klug . C. . Tanabe . K. . 2021 . Ammonoid soft tissue remains revealed by computed tomography . Swiss Journal of Palaeontology . 140 . 1 . Article 11 . 10.1186/s13358-021-00226-y . free . 11336/135617 . free .
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  52. Weis . R. . Dzyuba . O. S. . Dietze . V. . Riegraf . W. . A new belemnite species from the Aalenian of the Swabian Alb (SW Germany) and its position in the phylogeny of Megateuthididae (Belemnitida) . 2021 . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 300 . 1 . 23–31 . 10.1127/njgpa/2021/0977. 0077-7749 . 234855507 .
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  78. Chernyshev . A. V. . Goedert . J. L. . New species of latest Eocene/earliest Oligocene microgastropods (Heterobranchia: Orbitestellidae and Omalogyridae) from the Gries Ranch Formation, Lewis County, Washington State, USA . 2021 . Zootaxa . 4981 . 3 . 469–480 . 10.11646/zootaxa.4981.3.3 . 34186710 . 235686756 .
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