This list of fossil fish research presented in 2021 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2021.
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Falxcornus[1] | Gen. et sp. nov | In press | Meng & Gai | Devonian (Lochkovian) | Xishancun Formation | A member of Galeaspida belonging to the family Tridensaspidae. Genus includes new species F. liui. | ||
Hongshanaspis[2] | Gen. et sp. nov | Valid | Liu et al. | Silurian (Telychian) | Qingshui Formation | A member of Galeaspida belonging to the family Hanyangaspidae. Genus includes new species H. inexpectatus. | ||
Jiangxialepis[3] | Gen. et sp. nov | Valid | Liu et al. | Silurian (Telychian) | Fentou Formation | A member of Galeaspida belonging to the group Eugaleaspidiformes. The type species is J. retrospina. | ||
Qushiaspis[4] | Gen. et sp. nov | Valid | Jiang et al. | Early Devonian | Xujiachong Formation | A member of Galeaspida. Genus includes new species Q. elaia. | ||
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Bianchengichthys[13] | Gen. et sp. nov | Valid | Li, Zhu & Zhu in Li et al. | Silurian (Ludfordian) | Xiaoxi Formation | A placoderm closely related to the last common ancestor of bony and cartilaginous fishes. The type species is B. micros. | ||
Leptodontichthys[14] | Gen. et sp. nov | Jobbins et al. | Devonian (Givetian) | Taboumakhlouf Formation | A member of Arthrodira belonging to the family Plourdosteidae. The type species is L. ziregensis. | |||
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Dobunnacanthus[18] | Gen. et comb. nov | Valid | Dearden et al. | Devonian | A new genus for "Vernicomacanthus" waynensis Miles (1973) | |||
Nostolepis digitus[19] | Sp. nov | Valid | Li et al. | Devonian (Lochkovian) | Xitun Formation | |||
Nostolepis qujingensis | Sp. nov | Valid | Li et al. | Devonian (Lochkovian) | Xitun Formation | |||
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Aquilolamna[22] | Gen. et sp. nov | Vullo et al. | Late Cretaceous (Turonian) | Agua Nueva Formation | A probable planktivorous shark placed in the new family Aquilolamnidae, of uncertain placement. Possibly a member of Lamniformes. The type species is A. milarcae. | |||
Carcharhinus dudoni[23] | Sp. nov | Valid | Canevet & Lebrun | Miocene | A species of Carcharhinus. | |||
Carcharhinus pedronii | Sp. nov | Valid | Canevet & Lebrun | Miocene | A species of Carcharhinus. | |||
Carcharhinus tingae[24] | Sp. nov | Valid | Cicimurri & Ebersole | Eocene (Bartonian) | A species of Carcharhinus. | |||
Cladodus gailensis[25] | Sp. nov | Valid | Feichtinger et al. | Carboniferous (Serpukhovian) | ||||
Dracopristis[26] | Gen. et sp. nov | Valid | Hodnett et al. | Late Carboniferous (Kasimovian) | Atrasado Formation | A medium-sized ctenacanthiform shark known from a complete skeleton with soft tissue. The type species is D. hoffmanorum. | ||
Durnonovariaodus[27] | Gen. et sp. nov | Valid | Stumpf et al. | Late Jurassic (Tithonian) | Kimmeridge Clay | A member of the family Hybodontidae. The type species is D. maiseyi. | ||
Favusodus[28] | Gen. et sp. nov | Valid | Li et al. | Triassic (Ladinian–Carnian) | A member of Euselachii. Genus includes new species F. orientalis. | |||
Junggarensis[29] | Gen. et sp. nov | Valid | Roelofs et al. | Devonian (Famennian) | Genus includes new species J. ambiguus. | |||
Keichouodus | Gen. et sp. nov | Valid | Li et al. | Triassic (Ladinian–Carnian) | A member of Euselachii. Genus includes new species K. nimaiguensis. | |||
Keuperodus[30] | Gen. et comb. nov | Valid | Ivanov, Duffin & Richter | Late Triassic (Carnian) | Arden Sandstone Formation Grabfeld Formation | A member of the family Jalodontidae. Genus includes "Phoebodus" brodiei Woodward (1893) (interpreted by Ivanov, Duffin & Richter, 2021 as a senior synonym of "Phoebodus" keuperinus Seilacher, 1948). | ||
Lilamna[31] | Gen. et comb. nov | Valid | Greenfield | late Eocene or Cretaceous (uncertain) | A possible member of the family Pseudoscapanorhynchidae. The type species is "Archaeolamna" apophysata Li (1997). | |||
Maiseyacanthus[32] | Gen. et sp. nov | Valid | Bronson | Carboniferous (Mississippian) | Fayetteville Shale | A symmoriiform. The type species is M. ozarkanum. | ||
Maiseyodus[33] | Gen. et comb. nov | Valid | Long et al. | Devonian (Emsian) | Cravens Peak Beds | A member of the family Mcmurdodontidae; a new genus for "Mcmurdodus" whitei Turner & Young (1987). | ||
Moskovirhynchus[34] | Gen. et sp. nov | Valid | Popov & Shapovalov | Late Jurassic | A chimaera belonging to the family Callorhinchidae. Genus includes new species M. robustus. | |||
Nebriimimus[35] | Gen. et sp. nov | Valid | Collareta et al. | Pliocene (Zanclean) | A member of Rajiformes, possibly a skate. The type species is N. wardi. | |||
Negaprion cossardi | Sp. nov | Valid | Canevet & Lebrun | Miocene | A species of Negaprion. | |||
Phoebodus curvatus[36] | Sp. nov | Valid | Ivanov | Devonian (Givetian–Frasnian) | ||||
Protochimaera[37] | Gen. et sp. nov | Valid | Lebedev & Popov in Lebedev et al. | Carboniferous (Viséan–Serpukhovian) | Dashkovo Formation | A chimaera. Genus includes new species P. mirabilis. | ||
Pseudocorax kindlimanni[38] | Sp. nov | In press | Jambura, Stumpf & Kriwet | Late Cretaceous (Cenomanian) | Sannine Formation | |||
Ptychodus maghrebianus[39] | Sp. nov | In press | Amadori et al. | Late Cretaceous (Turonian) | ||||
Reifella[40] | Gen. et sp. nov | Valid | Ivanov in Ivanov et al. | Permian (Roadian) | Cutoff Formation | A member of the family Anachronistidae. Genus includes new species R. lata. | ||
Rosaodus | Gen. et sp. nov | Valid | Li et al. | Triassic (Ladinian–Carnian) | A member of Elasmobranchii of uncertain phylogenetic placement. Genus includes new species R. xingyiensis. | |||
Strophodus indicus[41] | Sp. nov | Valid | Sharma & Singh | Middle Jurassic (Bathonian) | Jaisalmer Formation | Marine hybodont shark | ||
Strophodus jaisalmerensis[42] | Sp. nov | In press | Kumar et al. | Jurassic | Jaisalmer Formation | A hybodont shark. | ||
Taeniurops sergiomorai[43] | Sp. nov | Laurito & Valerio | Miocene–Pliocene (Messinian–Piacenzian) | Uscari Formation | A stingray, a species of Taeniurops. | |||
Triodus aeduorum[44] | Sp. nov | Valid | Luccisano et al. | Early Permian | Autun Basin | |||
Vallisodus[45] | Nom. nov | Valid | Duffin | Late Triassic | Penarth Group | A member of Neoselachii; a replacement name for Vallisia Duffin (1982). | ||
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Achirus australis[67] | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Achirus. | |||
Achirus chungkuz | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Achirus. | |||
Agonopsis cume | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Agonopsis. | |||
Agoultpycnodus[68] | Gen. et comb. nov | Valid | Taverne & Capasso | Late Cretaceous (Cenomanian-Turonian) | Akrabou Formation | A member of the family Pycnodontidae. The type species is A. aldrovandii. | ||
Alilepis texasensis | Sp. nov | Valid | Bakaev in Ivanov et al. | Permian (Roadian) | Cutoff Formation | A member of the family Elonichthyidae. | ||
Anomoeodus aegypticus[69] | Sp. nov | Valid | Capasso et al. | Late Cretaceous (Maastrichtian) | Dakhla Formation | A member of Pycnodontiformes. | ||
Anomoeodus caddoi[70] | Sp. nov | Valid | Suarez et al. | Early Cretaceous (Albian) | Holly Creek Formation | A member of Pycnodontiformes. | ||
Aphanolebias bettinae[71] | Sp. nov | In press | Bradić-Milinović, Rundić & Schwarzhans | Miocene | A member of the family Valenciidae. | |||
Apuliadercetis gonzalezae[72] | Sp. nov | In press | Díaz-Cruz, Alvarado-Ortega & Cantalice | Late Cretaceous (Campanian) | Angostura Formation | A member of Aulopiformes belonging to the family Dercetidae. | ||
Archaemacruroides vanknippenbergi[73] | Sp. nov | In press | Schwarzhans & Jagt | Late Cretaceous (Maastrichtian) | Maastricht Formation | A member of Gadiformes of uncertain phylogenetic placement. | ||
Ariosoma mesohellenica[74] | Sp. nov | Valid | Agiadi, Koskeridou & Thivaiou | Miocene (Aquitanian) | A species of Ariosoma. | |||
Austelliscus[75] | Gen. et sp. nov | Figueroa, Weinschütz & Friedman | Middle Devonian or older | Paraná Basin | An early ray-finned fish. Genus includes new species A. ferox. | |||
Auxis koreanus[76] | Sp. nov | Valid | Nam, Nazarkin & Bannikov | Middle Miocene | Duho Formation | A species of Auxis. | ||
Bardackichthys[77] | Gen. et sp. nov | In press | Hacker & Shimada | Late Cretaceous (Cenomanian) | Woodbine Formation | A member of Ichthyodectiformes. Genus includes new species B. carteri. | ||
Bobbitichthys[78] | Gen. et comb. nov | Valid | Schwarzhans, Milàn & Carnevale | Paleocene (Selandian) | Kerteminde Marl | A member of the family Macrouridae. The type species is "Hymenocephalus" rosenkrantzi Schwarzhans (2003). | ||
Brauccipycnodus[79] | Gen. et comb. nov | Valid | Taverne & Capasso | Early Cretaceous (Albian) | A member of the family Pycnodontidae. The type species is "Proscinetes" pillae Capasso (2007). | |||
Briveichthys[80] | Gen. et sp. nov | Valid | Štamberg & Steyer | Permian | Brive Basin | A pygopterid. Genus includes new species B. chantepieorum. | ||
Capromimus undulatus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Capromimus. | |||
Cheirolepis jonesi[81] | Sp. nov | In press | Newman et al. | Devonian (Givetian) | Tordalen Formation | |||
Chiloconger chilensis | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Chiloconger. | |||
Choichix[82] | Gen. et sp. nov | Valid | Cantalice, Than-Marchese & Villalobos-Segura | Late Cretaceous (Cenomanian) | A member of Acanthopterygii of uncertain phylogenetic placement. Genus includes new species C. alvaradoi. | |||
Citharichthys parvisulcus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún beds Lacui Formation Navidad Formation Ranquil Formation | A species of Citharichthys. | |||
Citharichthys vergens | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Citharichthys. | |||
Clarotes eocenicus[83] | Sp. nov | Valid | Murray & Holmes | Late Eocene | Jebel Qatrani Formation | A species of Clarotes. | ||
Coccolepis solnhofensis[84] | Sp. nov | Valid | López-Arbarello & Ebert | Late Jurassic (Tithonian) | Altmühltal Formation | A member of Chondrostei belonging to the family Coccolepididae. | ||
Coelorinchus fidelis | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún Beds Lacui Formation Navidad Formation | A species of Coelorinchus. | |||
Coelorinchus rapelanus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Coelorinchus. | |||
Cottunculus primaevus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Cottunculus. | |||
Cretaserranus | Gen. et sp. nov | In press | Schwarzhans & Jagt | Late Cretaceous (Maastrichtian) | Maastricht Formation | A member of Perciformes, possibly belonging to the family Serranidae. Genus includes new species C. maastrichtensis. | ||
Ctenopharyngodon orientalis[85] | Sp. nov | Valid | Su, Chang & Chen | Miocene | Xiacaowan Formation | A species of Ctenopharyngodon. | ||
Ctenopharyngodon xiejiaensis | Sp. nov | Valid | Su, Chang & Chen | Miocene | Xiejia Formation | A species of Ctenopharyngodon. | ||
Dezaoia | Gen. et sp. nov | Valid | Su, Chang & Chen | Oligocene | Ulanbulage Formation | A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is D. saintjaquensis. | ||
Diastemapycnodus[86] | Gen. et sp. nov | In press | Abu El-Kheir et al. | Late Cretaceous (Maastrichtian) | Dakhla Formation | A member of Pycnodontiformes. Genus includes new species D. tavernensis. | ||
Elongofuro augustilgi[87] | Sp. nov | Valid | Ebert | Late Jurassic (Kimmeridgian) | Nusplingen Limestone | A member of Ophiopsiformes. | ||
Eoctenopharyngodon | Gen. et sp. nov | Valid | Su, Chang & Chen | Oligocene | Dongying Formation | A member of the family Cyprinidae belonging to the subfamily Squaliobarbinae. The type species is E. liui. | ||
Eotexachara[88] | Gen. et sp. nov | In press | Wick | Late Cretaceous (Campanian) | Aguja Formation | A member of Characiformes. Genus includes new species E. malateres. | ||
Feroxichthys panzhouensis[89] | Sp. nov | Valid | Ma, Xu & Geng | Middle Triassic (Anisian) | Guanling Formation | A member of the family Colobodontidae. | ||
Garganomyctophum[90] | Gen. et sp. nov | Valid | Taverne | Late Cretaceous (Santonian) | A lanternfish. The type species is G. sorbinii. | |||
Gnathophis elongatus | Sp. nov | Valid | Agiadi, Koskeridou & Thivaiou | Miocene (Aquitanian) | A species of Gnathophis. | |||
Gnathophis quinzioi | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Gnathophis. | |||
Gobiosoma? axsmithi[91] | Sp. nov | Valid | Ebersole, Cicimurri & Stringer | Oligocene (Rupelian) | Byram Formation | A member of the family Gobiidae. | ||
Guiclupea[92] | Gen. et sp. nov | Valid | Chen et al. | Oligocene | A member of Clupeomorpha belonging to the group Ellimmichthyiformes. The type species is G. superstes. | |||
Hellenigobius[93] | Gen. et sp. et comb. nov | Valid | Schwarzhans, Agiadi & Thivaiou | Miocene | ? | A member of the subfamily Gobionellinae. The type species is H. praeschismatus; genus also includes "Pomatoschistus" bunyatovi Bratishko et al. (2015). | ||
Katyagobius[94] | Gen. et sp. nov | Valid | Reichenbacher & Bannikov | Miocene | A member of the family Gobiidae. The type species is K. prikryli. | |||
Kelliaichthys[95] | Nom. nov | Valid | Schultze in Schultze et al. | A replacement name for Kellia Kazantseva-Selezneva (1980). | ||||
Klincigobus haraldahnelti | Sp. nov | In press | Bradić-Milinović, Rundić & Schwarzhans | Miocene | A member of the family Gobiidae. | |||
Krumvirichthys[96] | Gen. et sp. nov | Valid | Přikryl | Oligocene–early Miocene | A deep-sea smelt. The type species is K. brzobohatyi. | |||
Kuhlia orientalis | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A flagtail. | |||
Lampanyctus ipunensis | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún beds | A species of Lampanyctus. | |||
Larimichthys koae[97] | Sp. nov | In press | Lin & Chien | Late Miocene | Tapu Formation | A species of Larimichthys. | ||
Lepophidium chonorum | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Lepophidium. | |||
Lepophidium mapucheorum | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Lepophidium. | |||
Maurolicus brevirostris | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Maurolicus. | |||
Navidadichthys | Gen. et sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | Possibly a member of the family Prototroctidae. The type species is N. mirus. | |||
Neuburgichthys | Nom. nov | Valid | Schultze in Schultze et al. | A replacement name for Neuburgia Kazantseva-Selezneva (1980). | ||||
Nezumia epuge | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation Navidad Formation | A species of Nezumia. | |||
Nibea chaoi | Sp. nov | In press | Lin & Chien | Late Miocene | Tapu Formation | A species of Nibea. | ||
Oshiaichthys | Nom. nov | Valid | Schultze in Schultze et al. | Triassic | A replacement name for Oshia Sytchevskaya (1999). | |||
Palaeopantodon[98] | Gen. et sp. nov | Valid | Taverne | Late Cretaceous (Cenomanian) | A member of the family Pantodontidae. The type species is P. vandersypeni. | |||
Pankowskipiscis[99] | Gen. et sp. nov | Valid | Taverne | Late Cretaceous (Cenomanian) | A member of the family Pantodontidae. The type species is P. haqelensis. | |||
Paracarapus | Gen. et sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A pearlfish. The type species is P. chilensis. | |||
Peltoperleidus asiaticus[100] | Sp. nov | Valid | Xu | Middle Triassic (Anisian) | Guanling Formation | A member of Neopterygii belonging to the group Louwoichthyiformes. | ||
Petersichthys[101] | Gen. et sp. nov | Valid | Taverne | Late Cretaceous (Cenomanian) | A member of the family Pantodontidae. The type species is P. libanicus. | |||
Physiculus pichi | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Physiculus. | |||
Pinichthys shirvanensis[102] | Sp. nov | Valid | Bannikov | Miocene | A member of Perciformes belonging to the group Stromateoidei. | |||
Plesiogobius | Gen. et sp. et comb. nov | Valid | Schwarzhans, Agiadi & Thivaiou | Miocene (Aquitanian) | A member of the family Gobiidae belonging to the subfamily Gobiinae. The type species is P. felliensis. | |||
Polyipnus bandeli | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Polyipnus. | |||
Portentosoceros[103] | Gen. et comb. nov | Valid | Nazarkin & Bannikov | Miocene | A member of Percoidei of uncertain phylogenetic placement; a new genus for "Pentaceros" sakhaliniсus Gretchina (1975). | |||
Primuluchara | Gen. et sp. nov | In press | Wick | Late Cretaceous (Campanian) | Aguja Formation | A member of Characiformes. Genus includes new species P. laramidensis. | ||
Pseudohilsa nikosi[104] | Sp. nov | Valid | Kevrekidis, Arratia & Reichenbacher in Kevrekidis et al. | Late Miocene | A member of the family Clupeidae. | |||
Pseudolesueurigobius | Gen. et sp. nov | Valid | Reichenbacher & Bannikov | Miocene | A member of the family Gobiidae. The type species is P. manfredi. | |||
Pseudonus humilis | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Ipún beds Navidad Formation | A species of Pseudonus. | |||
Pteronisculus changae[105] | Sp. nov | Valid | Ren & Xu | Middle Triassic (Anisian) | Guanling Formation | |||
Pterothrissus transpacificus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Pterothrissus. | |||
Pythonichthys panulus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation | A species of Pythonichthys. | |||
Raususetarches[106] | Gen. et sp. nov | Valid | Yabumoto & Nazarkin | Late Miocene | Koshikawa Formation | A member of the family Scorpaenidae. Genus includes new species R. sakurai. | ||
Rhinocephalus cretaceus | Sp. nov | In press | Schwarzhans & Jagt | Late Cretaceous (Maastrichtian) | Maastricht Formation | A member of the family Merlucciidae. | ||
Rhynchoconger chiloensis | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Rhynchoconger. | |||
Sarmatigobius | Gen. et sp. et comb. nov | Valid | Reichenbacher & Bannikov | Miocene | A member of the family Gobiidae. The type species is S. compactus; genus also includes "Hesperichthys" iugosus Schwarzhans, Brzobohatý & Radwańska (2020). | |||
Saurichthys sceltrichensis[107] | Sp. nov | Valid | Renesto, Magnani & Stockar | Middle Triassic (Ladinian) | Meride Limestone | |||
Severnichthus[108] [109] | Gen. et sp. nov | In press | Stringer & Schwarzhans | Late Cretaceous (Maastrichtian) | Severn Formation | Possibly a member of Polymixiiformes. Genus includes new species S. bourdoni. | ||
Sirembola supersa | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Lacui Formation | A cusk-eel. | |||
Spectrunculus sparsus | Sp. nov | Schwarzhans & Nielsen | Early Miocene | Navidad Formation | A species of Spectrunculus. | |||
Taosciaena | Gen. et 2 sp. nov | In press | Lin & Chien | Late Miocene | Tapu Formation | A member of the family Sciaenidae. The type species is T. jiangi; genus also includes T. hui. | ||
Toxopyge vracevicensis | Sp. nov | In press | Bradić-Milinović, Rundić & Schwarzhans | Miocene | A member of the family Gobiidae. | |||
Wilsonium[110] | Gen. et comb. nov | Valid | Liu | Early Eocene | Allenby Formation | A Catostomidae sucker. The type species is "Amyzon" brevipinne (1893). | ||
Yarigobius | Gen. et 2 sp. nov | Valid | Reichenbacher & Bannikov | Miocene | A member of the family Gobiidae. The type species is Y. decoratus; genus also includes Y. naslavcensis. | |||
Name | Novelty | Status | Authors | Age | Type locality | Country | Notes | Images |
---|---|---|---|---|---|---|---|---|
Ceratodus guanganensis[128] | Sp. nov | Valid | Wang et al. | Late Jurassic | Shaximiao Formation | Announced in 2021; the final version of the article naming it was published in 2022. | ||
Eusthenodon bourdoni[129] | Sp. nov | Valid | Downs, Barbosa & Daeschler | Devonian (Famennian) | Catskill Formation | |||