2021 in paleoentomology explained

See also: 2021 in arthropod paleontology and 2021 in paleontology. This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

Clade Amphiesmenoptera

Lepidoptera

New Lepidopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acanthocorona^[1]	Gen. et 7 sp. nov	Valid	Mey, Léger & Lien	Cretaceous Cenomanian	Burmese amber		A Lophocoronidae moth. The type species is A. skalskii The genus also includes A. bowangi, A. kuranishii, A. muelleri, A. sattleri, A. spinifera, & A. wichardi.
Agathiphagama	Gen. et sp. nov	Valid	Mey, Léger & Lien	Cretaceous Cenomanian	Burmese amber		An agathiphagid aglossatan moth. The type species is A. perdita.
Phyllocnistis cretacea^[2]	Sp. nov	Valid	Fischer	Late Cretaceous (Cenomanian)	Burmese amber		A species of Phyllocnistis
Phyllonorycter inopinata	Sp. nov	Valid	Fischer	Lutetian	Baltic amber		A species of Phyllonorycter

Lepidopteran research

• The first known Cretaceous caterpillar armoured with spines is described from the Burmese amber by Haug & Haug (2021).^[3]
• Fossil leaf mine produced by a lyonetiid-like leaf-mining moth, representing the oldest record of such moths reported to date, is described from the Upper Cretaceous (Campanian) Kaiparowits Formation (Utah, United States) by Maccracken et al. (2021), who name a new ichnotaxon Leucopteropsa spiralae.^[4]

Trichopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Archaeopolycentra achupomotet^[5]	Sp. nov	Valid	Melnitsky & Ivanov	Late Cretaceous (Santonian)	Kheta Formation (Taymyr amber)		A member of the family Polycentropodidae.
Archaeopolycentra turvalsia	Sp. nov	Valid	Melnitsky & Ivanov	Late Cretaceous (Santonian)	Kheta Formation (Taymyr amber)		A member of the family Polycentropodidae.
Cernotina fossilinova^[6]	Sp. nov	Valid	Wichard & Neumann	Probably early Miocene	Dominican amber		A caddisfly, a species of Cernotina.
Electrotrichia rasnitsyni^[7]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Late Eocene	Rovno amber		A member of the family Hydroptilidae.
Folindusia aenigmatosa^[8]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Folindusia expansa	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Folindusia inconspicua	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Folindusia inventa	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Folindusia robusta	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Folindusia tortilis	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Holocentropus tutkaktut^[9]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Eocene (Priabonian)	Rovno amber		A species of Holocentropus.
Palaeopsilotreta kachini^[10]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber		A member of the family Odontoceridae.
Palaeopsilotreta succini	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber		A member of the family Odontoceridae.
Plectrocnemia kirmikhia^[11]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Eocene (Priabonian)	Rovno amber		A species of Plectrocnemia.
Psilotreta fossilis	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber		A species of Psilotreta.
Purbimodus khramovi^[12]	Sp. nov	Valid	Sukatsheva & Aristov	Early Cretaceous			A member of the family Vitimotauliidae.
Terrindusia nova	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous			A larval case of a caddisfly.
Wormaldia denticulata^[13]	Sp. nov	In press	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A caddisfly, a species of Wormaldia.
Wormaldia diplobifurca	Sp. nov	In press	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A caddisfly, a species of Wormaldia.

Clade Antliophora

Dipterans

New Dipteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acartophthalmites crassipes^[14]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber		A member of Opomyzoidea belonging to the new family Clusiomitidae.
Acartophthalmites luridus	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber		A member of Opomyzoidea belonging to the new family Clusiomitidae.
Acartophthalmites rugosus	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber		A member of Opomyzoidea belonging to the new family Clusiomitidae.
Aleksiana^[15]	Gen. et sp. et comb. nov	Valid	Krzemiński et al.	Cretaceous	Burmese amber		A member of the crane fly family Limoniidae. The type species is A. rasnitsyni; genus also includes "Antocha" lapra Podenas & Poinar (2009).
Allarete patriciae^[16]	Sp. nov	Valid	Nel	Early Eocene	Oise amber		A member of the family Cecidomyiidae.
Allodia paleoafricana^[17]	Sp. nov	Valid	Bouju et al.	Early Miocene	Ethiopian amber		A species of Allodia.
Apocephalus chiapanecus^[18]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Mexican amber		A species of Apocephalus.
Apocephalus dominicanus	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Dominican amber		A species of Apocephalus.
Apocephalus miocenus	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene			A species of Apocephalus.
Araripogon pulchrus^[19]	Sp. nov	In press	Lamas, Sampronha & Ribeiro	Early Cretaceous (Aptian)	Crato Formation		A member of the family Asilidae.
Bashkonia^[20]	Gen. et sp. nov	Valid	Lukashevich	Middle Triassic	Röt Formation		A member of the family Nadipteridae. Genus includes new species B. franconica.
Bibio castaneipennis^[21]	Sp. nov	Valid	Skartveit & Wedmann	Oligocene			A species of Bibio.
Bibio succineus^[22]	Sp. nov	Valid	Skartveit	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Bibio.
Burmahesperinus^[23]	Gen. et 3 sp. nov	Valid	Ševčík et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Bibionidae. The type species is B. antennatus; genus also includes B. conicus and B. pedicellatus.
Burmasymmerus^[24]	Gen. et 2 sp. nov	Valid	Ševčík, Hippa & Burdíková	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Ditomyiidae. The type species is B. korneliae; genus also includes B. wieslawi.
Burmatricha	Gen. et sp. nov	Valid	Ševčík, Hippa & Burdíková	Late Cretaceous (Cenomanian)	Burmese amber		A member of Sciaroidea of uncertain phylogenetic placement. The type species is B. mesozoica.
Cascomixticus^[25]	Gen. et sp. nov	Valid	Poinar & Vega	Cretaceous (Albian/Cenomanian)	Burmese amber		A member of the family Apsilocephalidae. The type species is C. tubuliferous.
Clusiomites	Fam. et gen. et comb. et sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber		A member of Opomyzoidea, the type genus of the new family Clusiomitidae. The type species is "Acartophthalmites" clusioides Roháček (2016); genus also includes new species C. ornatus.
Crenoptychoptera gaoi^[26]	Sp. nov	Valid	Liu & Huang	Middle–Late Jurassic	Jiyuan Basin		A member of the family Ptychopteridae.
Cretobibio rasnitsyni^[27]	Sp. nov	Valid	Lukashevich, Amorim & Ribeiro	Early Cretaceous (Aptian)	Crato Formation		A member of the family Bibionidae.
Cretolimonia dayana^[28]	Sp. nov		Kopeć in Kopeć et al.	Jurassic/Cretaceous boundary	Glushkovo Formation		A member of the subfamily Architipulinae.
Cretolimonia lukashevichae	Sp. nov		Kopeć & Krzemiński in Kopeć et al.	Jurassic/Cretaceous boundary	Dain Formation		A member of the subfamily Architipulinae.
Cretolimonia mikolajczyki	Sp. nov		Kopeć, Krzemiński, Soszyńska-Maj in Kopeć et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the subfamily Architipulinae.
Cretolimonia pseudojurassica	Sp. nov		Krzemiński in Kopeć et al.	Jurassic/Cretaceous boundary	Dain Formation		A member of the subfamily Architipulinae.
Cretothereva^[29]	Gen. et sp. nov	In press	Carmo, Lamas & Ribeiro	Early Cretaceous	Crato Formation		A member of the family Therevidae. Genus includes new species C. antiqua.
Cretpenthetria^[30]	Gen. et sp. nov	Valid	Li, Zhang & Xiao	Cretaceous	Burmese amber		A member of the family Bibionidae. Genus includes new species C. burmensis.
Culex ekaterinae^[31]	Sp. nov	Valid	Giłka, Harbach & Perkovsky	Eocene	Rovno amber		A mosquito, a species of Culex.
Culicoides paleopestis^[32]	Sp. nov	In press	Peñalver et al.	Eocene	Anglesea amber		A species of Culicoides.
Cyttaromyia freiwaldi^[33]	Sp. nov	Valid	Kania-Kłosok et al.	Eocene	Green River Formation		A member of the family Cylindrotomidae.
Cyttaromyia gelhausi	Sp. nov	Valid	Kania-Kłosok et al.	Eocene	Green River Formation		A member of the family Cylindrotomidae.
Decessia^[34]	Gen. et sp. nov	Valid	Krzemiński et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the crane fly family Limoniidae. The type species is D. podenasi.
Elephantomyia christelae^[35]	Sp. nov		Kania-Kłosok & Krzemiński	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Elephantomyia.
Elephantomyia prima	Sp. nov		Kania-Kłosok & Krzemiński	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Elephantomyia.
Forcipomyia rasnitsyni^[36]	Sp. nov	Valid	Szadziewski, Sontag & Pankowski	Miocene	Ethiopian amber		A species of Forcipomyia.
Helius hispanicus^[37]	Sp. nov		Kania-Kłosok, Krzemiński & Arillo	Early Cretaceous (Albian)	Utrillas Group (Álava amber)		A species of Helius.
Helius turolensis	Sp. nov		Kania-Kłosok, Krzemiński & Arillo	Early Cretaceous (Albian)	Utrillas Group (San Just amber)		A species of Helius.
Hirmoneura messelense^[38]	Sp. nov	Valid	Wedmann et al.	Eocene (Lutetian)	Messel Formation		A member of the family Nemestrinidae.
Lebambromyia sacculifera^[39]	Sp. nov	Valid	Badano, Zhang, Yeates & Cerretti in Badano et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of Phoroidea.
Leptotarsus andradei^[40]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation		A species of Leptotarsus.
Leptotarsus gelhausi	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation		A species of Leptotarsus.
Leptotarsus ivoneae	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation		A species of Leptotarsus.
Leptotarsus lemeae	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation		A species of Leptotarsus.
Mesoacentron^[41]	Gen. et sp. nov	Valid	Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.	Late Cretaceous (Santonian)	Kheta Formation (Taimyr amber)		A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is M. kaluginae.
Meunierohelea anglesensis	Sp. nov	In press	Peñalver et al.	Eocene	Anglesea amber		A member of the family Ceratopogonidae.
Minyohelea nexuosa^[42]	Sp. nov	In press	Pielowska-Ceranowska, Azar & Szwedo	Early Cretaceous (Barremian)	Lebanese amber		A member of the family Ceratopogonidae.
Myriophora asiatica	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Zhangpu amber		A species of Myriophora.
Neoarisemus groehni^[43]	Sp. nov	Valid	Wagner	Eocene			A member of the family Psychodidae.
Orimarguloides^[44]	Gen. et sp. nov	Valid	Men & Podenas	Cretaceous	Burmese amber		A member of the crane fly family Limoniidae. Genus includes new species O. simplex.
Palaeocentron	Gen. et sp. nov	Valid	Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.	Cretaceous (probably Albian–Cenomanian)	Burmese amber		A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is P. krzeminskii.
Palaeoglaesum pilosus^[45]	Sp. nov	Valid	Skibińska, Krzemiński & Zhang in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Palaeoglaesum stebneri	Sp. nov	Valid	Skibińska & Krzemiński in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Palaeoglaesum teres	Sp. nov	Valid	Skibińska & Albrycht in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Paleoplatyura agnieszkae^[46]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber		A species of Paleoplatyura.
Paleoplatyura magnifica	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber		A species of Paleoplatyura.
Paleoplatyura miae	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber		A species of Paleoplatyura.
Paleotelmatoscopus^[47]	Gen. et comb. et sp. nov	Valid	Curler & Skibińska	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Psychodidae. The type species is "Pericoma" formosa Meunier (1905); genus also includes new species P. madrizi.
Polymera alexanderi^[48]	Sp. nov	Valid	Krzemiński et al.	Miocene	Dominican amber		A species of Polymera.
Protopenthetria^[49]	Gen. et sp. nov	Valid	Li et al.	Cretaceous	Burmese amber		A member of the family Bibionidae. Genus includes new species P. skartveiti.
Rhabdomastix asiatica^[50]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber		A species of Rhabdomastix.
Rhabdomastix cretacica	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber		A species of Rhabdomastix.
Rhabdomastix krzeminskae	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber		A species of Rhabdomastix.
Rhabdomastix myanmae	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber		A species of Rhabdomastix.
Rhipidoxylomyia rasnitsyni	Sp. nov	Valid	Nel	Early Eocene	Oise amber		A member of the family Cecidomyiidae.
Stempellinella gilkai^[51]	Sp. nov	Valid	Zakrzewska & Jankowska	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Stempellinella.
Trichoneura xavieri^[52]	Sp. nov	Valid	Kania-Kłosok et al.	Early Cretaceous (Albian)	Utrillas Group		A species of Trichoneura.
Zhangsolva burmensis^[53]	Sp. nov	In press	Zhang & Zhang	Cretaceous	Burmese amber		A member of the family Zhangsolvidae.

Dipteran research

• Baranov et al. (2021) document a high diversity of fly larvae from the Eocene Baltic amber, preserving taxonomically, morphologically, and ecologically important characters, and evaluate the implications of this finding for the knowledge of the functional ecology of the Eocene fly larvae.^[54]

Mecopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Conicholcorpa longa^[55]	Sp. nov		Zhang, Shih & Ren in Zhang et al.	Middle Jurassic	Jiulongshan Formation		A scorpionfly belonging to the family Holcorpidae.
Glanomerope^[56]	Gen. et sp. nov	Valid	Poschmann & Nel	Permian	Meisenheim Formation		A scorpionfly belonging to the family Protomeropidae. Genus includes new species G. virgoferroa.
Longiphlebia incompleta^[57]	Sp. nov	In press	Lian, Cai & Huang	Late Jurassic	Tiaojishan Formation		A scorpionfly belonging to the family Orthophlebiidae.
Mesopsyche jinsuoguanensis^[58]	Sp. nov	Valid	Lian, Cai & Huang	Middle Triassic	Tongchuan Formation		A member of the family Mesopsychidae. Originally described as a species of Mesopsyche; transferred by Lian et al. (2024) to the genus Turbidapsyche.^[59]
Mesopsyche liaoi	Sp. nov	Valid	Lian, Cai & Huang	Late Triassic	Huangshanjie Formation		A member of the family Mesopsychidae. Originally described as a species of Mesopsyche; transferred by Lian et al. (2024) to the genus Turbidapsyche.

Clade Archaeorthoptera

†Caloneurodea

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aviobiella^[60]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)			A member of Archaeorthoptera belonging to the order Caloneurodea. Genus includes new species A. garroustei.
Oudardgramma^[61]	Gen. et sp. nov		Nel & Roques	Carboniferous (Bashkirian)			The type species is O. bruayensis.
Paraspiculum	Gen. et sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)			A member of Archaeorthoptera belonging to the group Caloneurodea. Genus includes new species P. brauckmannii.

†Cnemidolestodea

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Avionxixia	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)			A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is A. gui.
Ctenoptilus frequens^[62]	Sp. nov	Valid	Chen et al.	Carboniferous (Bashkirian to Moscovian)	Yanghugou Formation		A member of Archaeorthoptera belonging to the family Ctenoptilidae.
Palatinarkema^[63]	Gen. et sp. nov	Valid	Nel & Poschmann	Early Permian (probably Asselian–?Sakmarian)	Meisenheim Formation		A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is P. prokopi.
Piesbergopterum	Gen. et 2 sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)			A member of Archaeorthoptera belonging to the group Cnemidolestodea. Genus includes new species P. punctatum Dvořák et al. and P. avionensis Nel & Roques.^[64]

Orthopterans

New Orthopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Amberotridactylus^[65]	Gen. et sp. nov	In press	Du, Xu & Zhang	Cretaceous	Burmese amber		A member of the family Tridactylidae. Genus includes new species A. cheni.
Archaboilus polyneurus^[66]	Sp. nov	Valid	Gu, Yue & Ren in Gu et al.	Middle Jurassic (Bathonian–Callovian)	Jiulongshan Formation		An ensiferan belonging to the group Hagloidea and the family Haglidae.
Caelielca^[67]	Gen. et sp. nov	In press	Uchida	Cretaceous	Burmese amber		A member of the family Elcanidae. Genus includes new species C. spinocrus.
Cascogryllus setosus	Sp. nov	In press	Du, Xu & Zhang	Cretaceous	Burmese amber		A member of the family Tridactylidae.
Cratoelcana rasnitsyni^[68]	Sp. nov	In press	Nel & Jouault	Early Cretaceous	Crato Formation		A member of the family Elcanidae.
Hukawnelca	Gen. et sp. nov	In press	Uchida	Cretaceous	Burmese amber		A member of the family Elcanidae. Genus includes new species H. gracile.
Panorpidium spica^[69]	Sp. nov	In press	Kim et al.	Early Cretaceous (Albian)	Jinju Formation		An orthopteran belonging to the family Elcanidae.
Permohagla^[70]	Gen. et sp. nov	Valid	Gorochov	Late Permian			A member of Hagloidea. Genus includes new species P. tomica.
Phyllotridactylus^[71]	Gen. et sp. nov	In press	Xu et al.	Cretaceous	Burmese amber		A member of the family Tridactylidae. Genus includes new species P. wangi.
Pseudoacrida sennlaubi	Sp. nov	In press	Nel & Jouault	Early Cretaceous	Crato Formation		A member of the family Locustopsidae.

Orthopteran research

• Revision of the fossil record of the family Trigonidiidae is published by Desutter-Grandcolas et al. (2021).^[72]

Other archaeorthopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Archaeogerarus^[74]	Fam. et gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)			A member of Archaeorthoptera belonging to the new family Archaeogeraridae. The type species is A. schubneli.
Avionugonioneura^[75]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)			A member of Archaeorthoptera possibly belonging to the family Nugonioneuridae. Genus includes new species A. jouaulti.
Contracladus^[76]	Gen. et sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)			A member of Archaeorthoptera of uncertain phylogenetic placement. Genus includes new species C. impar.
Duquesnia^[77]	Gen. et sp. nov	Valid	Nel & Roques in Nel, Roques & Duquesne	Carboniferous (Moscovian)			A member of Archaeorthoptera. Genus includes new species D. gallica.
Penanoptera^[78]	Gen. et sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin		A member of Archaeorthoptera. Genus includes new species P. manseri.

Clade Coleopterida

Coleoptera

Adephaga

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Antephilorhizus^[79]	Gen. et sp. nov	Valid	Kirichenko-Babko, Perkovsky & Vasilenko	Eocene	Rovno amber		A ground beetle belonging to the tribe Lebiini. Genus includes new species A. pripiatiensis.
Balticeler^[80]	Gen. et sp. nov	Valid	Schmidt & Maddison in Schmidt, Scholz & Maddison	Eocene	Baltic amber	Europe (Baltic Sea region)	A ground beetle belonging to the subfamily Trechinae. The type species is B. kerneggeri.
Calosoma kimi^[81]	Sp. nov	Valid	Lee & Nam	Early Miocene	Geumgwangdong Formation		A species of Calosoma.

Archostemata

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Jurodes shef^[82]	Sp. nov	Valid	Yan, Strelnikova & Cai	Early Cretaceous (Berriasian-Barremian)	Gusinoe Ozero Group		A Jurodid beetle.
Kirejtomma^[83]	Gen. et comb. nov	Valid	Li & Cai	Cretaceous	Burmese amber		An Ommatid beetle The type species is "Clessidromma" zengi (2020).
Limnomma^[84]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Middle Jurassic	Haifanggou Formation		An Ommatid beetle. The type species is L. daohugouense.
Omma axsmithi^[85]	Sp. nov	In press	Jarzembowski, Zheng & Zhao	Cretaceous	Burmese amber		An Omma species Ommatid beetle.
Omma forte^[86]	Sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Albian to Cenomanian)	Burmese amber		An Omma species Ommatid beetle.
Paraodontomma leptocristatum^[87]	Sp. nov	Valid	Li, Yamamoto & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber		An Ommatid beetle.
Permocatinus^[88]	Gen. et sp. nov	Valid	Ponomarenko	Late Permian			A Coleocatiniid beetle. The type species is P. tomiensis.
Tomiaplus	Gen. et comb. et sp. nov	Valid	Ponomarenko	Late Permian			A Triaplid beetle. The type species is "Triaplus" sibiricus (2013), as well as new species T. minimus.
Tunguskagyrus уani	Sp. nov	Valid	Ponomarenko	Late Permian			A Triaplid beetle.
Zygadenia alexrasnitsyni^[89]	Sp. nov	Valid	Strelnikova & Yan	Early Cretaceous	Zaza Formation		An Ommatid beetle.

Polyphaga

Bostrichiformia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Archaenosodendron^[91]	Gen. comb. et 3 sp. nov	Valid	Li & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber		A nosodendrid beetle. The type species is "Nosodendron" cretaceum; genus also includes new species A. angulare, A. explanatum and A. remotidens.
Cretomegatoma^[92]	Gen. et comb. nov	Valid	Háva	Late Cretaceous (Cenomanian)	Burmese amber		A carpet beetle. The type species is "Megatoma" atypica (2017).
Mesonosa^[93]	Gen. et sp. nov	Valid	Tihelka et al.	Late Cretaceous (Cenomanian)	Burmese amber		A wounded-tree beetle. The type species is M. scandens.
Petalium bruteno^[94]	Sp. nov	Valid	Alekseev & Bukejs	Late Eocene Priabonian	Baltic amber		A Petalium species death watch beetle.
Petalium widewuto	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber		A Petalium species death watch beetle.
Trogoderma ainu^[95]	Sp. nov	Valid	Perkovsky, Háva & Zaitsev	Middle Eocene	Sakhalin amber		A Trogoderma species carpet beetle.
Tuberphradonoma	Gen. et sp. nov	Valid	Háva	Late Cretaceous Cenomanian)	Burmese amber		A carpet beetle. The type species is T. burmitica.
Xyletinus lobanovi^[96]	Sp. nov	Valid	Bukejs, Alekseev & Háva	Eocene	Baltic amber		A Xyletinusspecies death watch beetle.

=New Cucujiformian taxa

=

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aberrokorynetes oceanojubilaei^[97]	Sp. nov	In press	Kolibáč et al.	Eocene	Baltic amber		A Korynetinae checkered beetle.
Aliattalus^[98]	Gen. et sp. nov	Valid	Tshernyshev	Late Eocene	Baltic amber		A Malachiinae malachite beetle. The type species is A. intercalaris.
Archaeosciaphilus gyrommatus^[99]	Sp. nov	In press	Legalov & Poinar	Eocene	Baltic amber		A Polydrusini Entiminae broad nosed weevil.
Arkagalarhinus^[100]	Gen. et sp. nov	In press	Legalov	Late Cretaceous (Santonian-Campanian)	Arkagala Formation		A Chilecarinae ithycerid weevil. The type species is A. zherikhini.
Asiomira dubrovinae^[101]	Sp. nov	Valid	Nabozhenko & Bukejs	Eocene	Baltic amber		An Alleculinae darkling beetle.
Aulacoscelis santanensis^[102]	Sp. nov	In press	Legalov	Early Cretaceous (Aptian-Albian)	Santana Group		An orsodacnid leaf beetle.
Baltocar sontagae^[103]	Sp. nov	Valid	Bukejs & Legalov	Eocene	Baltic amber		A Sayrevilleinae tooth-nosed snout weevil.
Baltosidis^[104]	Gen. et 3 sp. nov	Valid	Szawaryn	Eocene	Baltic amber		A Microweiseini Microweiseinae lady beetle. The type species is B. damzeni; genus also includes B. damgaardi and B. szadziewskii.
Bronchotibia^[105]	Gen. et sp. nov	In press	Poinar & Legalov	Miocene (Burdigalian)	Dominican amber		A Lymantini Molytinae weevil. The type species is B. adunatus.
Cathartosilvanus perkovskyi^[106]	Sp. nov	Valid	Alekseev & Bukejs	Late Eocene	Rovno amber		A silvanid beetle.
Ceutorhynchus zerovae^[107]	Sp. nov	Valid	Legalov et al.	Late Eocene	Rovno amber		A weevil.
Coslonatus^[108]	Gen. et sp. nov	Valid	Li & Cai	Cenomanian	Burmese amber		A Colydiinae? Zopheridae tenebrionoid beetle. The type species is C. rasnitsyni.
Cretaidgia^[109]	Gen. et sp. nov	In press	Zhao, Liu & Yu	Late Cretaceous (Cenomanian)	Burmese amber		A prionocerid beetle. The type species is C. burmensis.
Diapus ethiopicus^[110]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Ethiopian amber		A Platypodinae pinhole borer ambrosia weevil.
Diapus resinae	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Zhangpu amber		A Platypodinae pinhole borer ambrosia weevil.
Dorytomus mikhailovi^[111]	Sp. nov	Valid	Legalov, Nazarenko & Perkovsky	Late Eocene	Rovno amber		A species of Dorytomus Curculioninae weevil.
Electroxylita^[112]	Gen. et sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber		A false darkling beetle. The type species is E. chronographica.
Eurocrossotus^[113]	Gen. et sp. nov	Valid	Vitali	Eocene	Baltic amber	Europe (Baltic Sea region)	A longhorn beetle belonging to the tribe Crossotini. The type species is E. alekseevi.
Glaesotropis rohdendorfi^[114]	Sp. nov	Valid	Legalov, Nazarenko & Perkovsky	Eocene (Priabonian)	Rovno amber		A zygaenodinin anthribine anthribid fungus weevil.
Glesoconomorphus ekaterinae^[115]	Sp. nov	Valid	Telnov et al.	Late Eocene Priabonian	Rovno amber		A mycterid palm beetle.
Gracilenticrus^[116]	Gen. et sp. nov	Valid	Yu, Kolibáč & Ślipiński in Yu et al.	Late Cretaceous (Cenomanian)	Burmese amber		A lophocaterid beetle. The type species is G. burmiticus.
Heterhelus buzina^[117]	Sp. nov	Valid	Kupryjanowicz, Lyubarsky & Perkovsky	Eocene	Rovno amber		A Kateretid short-winged flower beetle. Originally described as a species of Heterhelus, but Kirejtshuk, Jenkins Shaw & Smirnov (2023) transferred it to the genus Antirhelus.^[118]
Isomira lobanovi^[119]	Sp. nov	Valid	Nabozhenko & Bukejs	Eocene	Baltic amber		An Isomira species claw comb darkling beetle.
Kaszubia^[120]	Gen. et sp. nov	Valid	Legalov, Kupryjanowicz & Perkovsky	Late Eocene Priabonian	Baltic amber		A cossonine weevil. The type species is K. wanati.
Kulindrobor^[121]	Gen. et 2 sp. nov	Valid	Tihelka et al.	Late Cretaceous Cenomanian	Burmese amber		A tenebrionoid beetle of uncertain phylogenetic placement. Genus includes new species K. enigmaticus and K. magnus.
Lobanoviella^[122]	Gen. et sp. nov	In press	Kirejtshuk & Reid	Eocene	Baltic amber		A lobanovielline palophagine megalopodid leaf beetle. The type species is L. andreyi.
Madelinia capillata	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber		A false darkling beetle.
Melanosiagon^[123]	Gen. et sp. nov	Valid	Batelka & Prokop	Late Cretaceous (Cenomanian)	Burmese amber		A wedge-shaped beetle. The type species is M. serraticornis.
Metaclisa ottoi^[124]	Sp. nov	Valid	Nabozhenko, McKellar & Bukejs	Late Eocene Priabonian	Baltic amber		AMetaclisa species metaclisine darkling beetle.
Microscapha kugelanni	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber		A false darkling beetle.
Microtrogossita^[125]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous Cenomanian	Burmese amber		A Trogossitid bark-gnawing beetle. The type species is M. qizhihaoi.
Monolepta rappsilberi^[126]	Sp. nov	Valid	Bukejs et al.	Late Eocene Priabonian	Bitterfeld amber		A Monolepta species skeletonizing leaf beetle.
Neolatitergum^[127]	Nom. nov	Valid	Zhang	Jurassic	Jiulongshan Formation		A Trogossitid bark-gnawing beetle. A replacement name for "Latitergum" Yu et al. (2014).
Oedemera girulskii^[128]	Sp. nov	Valid	Szawaryn, Sontag & Kubisz	Late Eocene Priabonian	Baltic amber		An Oedemera species false blister beetle.
Pacyclaxyra^[129]	Gen. et sp. nov	Valid	Tihelka, Huang & Cai	Late Cretaceous Cenomanian	Burmese amber		A sooty mould beetle. The type species is P. azari.
Palaeatalasis^[130]	Gen. et sp. nov	Valid	Legalov	Eocene (Ypresian)	Green River Formation		A Megamerini Sagrinae leaf beetle. The type species is P. monrosi.
Palaeoanoplus^[131]	Trib., Gen., et sp. nov	Valid	Legalov	Late Eocene Priabonian	Baltic amber		A Palaeoanoplini Curculioninae weevil. The type species is P. horridus.
Paleopegorhinus^[132]	Gen. et sp. nov	Valid	Poinar, Brown & Legalov	Late Cretaceous (Cenomanian)	Burmese amber		A Chilecarinae new york weevil. The type species is P. micrommatus.
Palaeopeplus^[133]	Gen. et sp. nov	Valid	Powell & Cline	Miocene (Burdigalina)	Dominican amber		A Cillaeinae sap beetle. The type species is P. cascus.
Pelretes^[134] ^[135] ^[136]	Gen. et sp. nov	Valid	Tihelka et al.	Cretaceous	Burmese amber		A short-winged flower beetle. The type species is P. vivificus .
Phloiotrya inmarinata	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber		A Phloiotrya species false darkling beetle.
Protodasytes^[137]	Gen. et sp. nov	Valid	Tihelka et al.	Late Cretaceous (Cenomanian)	Charentese amber		A soft-wing flower beetle. The type species is P. cretaceus.
Protomauroania^[138]	Gen. et 2 sp. nov	Valid	Tshernyshev in Tshernyshev & Shcherbakov	Late Eocene	Baltic amber Rovno amber	Baltic Sea region	A Dasytinae malachite beetle. The type species P. chaetophorata Genus also includes P. mikhailovi.^[139]
Protostomopsis^[140]	Gen. et sp. nov	Valid	Bukejs, Ślipiński & Alekseev in Bukejs et al.	Eocene	Baltic amber		A cerylonid beetle. The type species is P. pandema.
Pseudomesauletes lobanovi	Sp. nov	Valid	Bukejs & Legalov	Eocene	Baltic amber		A Rhynchitinae tooth-nosed snout weevil.
Pulchritudo^[141]	Gen. et sp. nov	Valid	Krell & Vitali	Eocene (Ypresian)	Green River Formation		A Sagrinae leaf beetle. The type species is P. attenboroughi.
Rhamphophorus^[142]	Trib., gen. et sp. nov	Valid	Poinar & Brown	Cretaceous Cenomanian	Burmese amber		A Rhamphophorini Cimberidinae pine flower weevil. The type species is R. legalovii.
Salignacicola	Gen. et comb. nov	Valid	Batelka & Prokop	Late Cretaceous (Cenomanian)			A wedge shaped beetle. The type species is "Macrosiagon" ebboi (2004).
Symphora glaesonauta	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber		A false darkling beetle.
Symphora pollocki	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber		A false darkling beetle.
Teredolaemus primigenius^[143]	Sp. nov	Valid	Alekseev et al.	Late Eocene Priabonian	Baltic amber		A teredid beetle.
Toxorhynchus michalskii	Sp. nov	Valid	Legalov	Late Eocene Priabonian	Baltic amber		An Apioninae straight-snouted weevil.
Trematosphindus^[144]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous (Cenomanian)	Burmese amber		A slime mold beetle. The type species is T. newtoni
Zaiwa^[145]	Gen. et sp. nov	Valid	Lyubarsky et al.	Cretaceous Cenomanian	Burmese amber		A lophocaterid beetle. The type species is Z. pankowskiorum.
Zherikhiniellus^[146]	Gen. et sp. nov	Valid	Legalov	late Eocene Priabonian	Baltic amber		A Nanophyinae straight-snouted weevil. The type species is Z. rarus.

Cucujiformia research

• A piece of Burmese amber preserving the complete series of free-living stages (except of the last larval instar) of a Cretaceous parasitoid beetle Paleoripiphorus is described by Batelka, Engel & Prokop (2021).^[147]

=New Elateriformian taxa

=

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Amberophytum maculatum^[148]	Sp. nov	Valid	Poinar, Vega & Legalov	Late Cretaceous (Cenomanian)	Burmese amber		A cerophytid rare click beetle.
Anoeuma^[149]	Gen. et sp. nov	Valid	Li, Kundrata & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber		An elateroid click beetle of uncertain phylogenetic placement. The type species is A. lawrencei.
Arturmiles^[150]	Gen. et sp. nov	Valid	Fanti	Eocene	Prussian Formation (Baltic amber)		A soldier beetle belonging to the tribe Cantharini. The type species is A. pankowskiarum.
Balistica vetai^[151]	Sp. nov	Valid	Muona	Eocene	Baltic amber		A Dirhagini Melasinae false click beetle.
Baltodascillus^[152]	Gen. et sp. nov	Valid	Kundrata et al.	Eocene	Baltic amber		A dascillid beetle. The type species is B. serraticornis.
Burmagrilus^[153]	Gen. et sp. nov	Valid	Jiang, Song & Wang in Jiang et al.	Cretaceous	Burmese amber		A metallic wood-boring beetle. The type species is B. cretacus.
Burmomiles bilineatimaculatus^[154]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber		A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.^[155]
Burmomiles laticollis	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber		A soldier beetle.
Burmomiles oblongoculus	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber		A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.
Captopus^[156]	Gen. et sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber		A throscid beetle. The type species is C. depressiceps.
Chelonarium andabata^[157]	Sp. nov	Valid	Alekseev & Bukejs in Alekseev et al.	Eocene	Baltic amber		A turtle beetle.
Chelonarium dingansich	Sp. nov	Valid	Alekseev & Bukejs in Alekseev et al.	Eocene	Baltic amber		A turtle beetle.
Chelonarium dominicanum^[158]	Sp. nov	Valid	Poinar & Háva	Burdigalian	Dominican amber		A turtle beetle.
Coomanius balticus	Sp. nov	Valid	Muona	Eocene	Baltic amber		A Nematodini Macraulacinae false click beetle.
Cretocantharis^[159]	Gen. et sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber		A soldier beetle. The type species is C. veda.
Cretopachyderes^[160]	Gen. et sp. nov	Valid	Kundrata et al.	Cretaceous Cenomanian	Burmese amber		An Agrypninae click beetle. The type species is C. burmitinus.
Cretophengodes^[161]	Fam. nov., Gen. et sp. nov	Valid	Li, Kundrata, Tihelka & Cai in Li et al.	Cretaceous (Albian or Cenomanian)	Burmese amber		A cretophengodid elateroid beetle. The type species is C. azari.
Dyscharachthis frickae	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe (Baltic Sea region)	A Dyscharachthini Eucneminae false click beetle.
Electrothroscus	Gen. et sp. nov	Valid	Li & Cai	Cretaceous Cenomanian	Burmese amber		A throscid false click beetle. The type species is E. yanpingae.
Entomophthalmus antennalis	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe (Baltic Sea region)	A Dirhagini Melasinae false click beetle.
Erdaia bispinulosa	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Orodotini? Macraulacinae false click beetle.
Erdaia longicornis	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Orodotini? Macraulacinae false click beetle.
Erdaia longitarsis	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Orodotini? Macraulacinae false click beetle.
Euryptychoides	Gen. et sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Euryptychini Macraulacinae false click beetle. The type species is E. magnus.
Euryptychus decoratus	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Euryptychini Macraulacinae false click beetle.
Euryptychus excavatus	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Euryptychini Macraulacinae false click beetle.
Euryptychus helmwigae	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Euryptychini Macraulacinae false click beetle.
Euryptychus sieglindae	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Euryptychini Macraulacinae false click beetle.
Euryptychus waltrautae	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		An Euryptychini Macraulacinae false click beetle.
Hukawngichthyurus maha	Sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber		A soldier beetle.
Isorhipis balticus	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		An Isorhipis species melasine false click beetle
Lycocerus crassepunctatus^[162]	Sp. nov	Valid	Kazantsev	Late Eocene Priabonian	Baltic amber		A soldier beetle.
Malthodes aliger	Sp. nov	Valid	Kazantsev	Late Eocene Priabonian	Baltic amber		A soldier beetle
Malthodes headsi^[163]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber		A soldier beetle.
Malthodes jaredi	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber		A soldier beetle, a species of Malthodes.
Malthodes vladiceniensis^[164]	Sp. nov	Valid	Pintilioaie, Fanti & Ionesi	Middle Miocene	Bârnova-Muntele Formation		A soldier beetle.
Microrhagus (Emyirhagus) vetai	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		A Microrhagusspecies melasine false click beetle
Murcybolus^[165]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous Cenomanian	Burmese amber		A Burmolycini dexorine net-winged beetle. The type species is M. longiantennus.
Nematodes schwertleitae	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		A Nematodini Macraulacinae false click beetle.
Neocharoides	Gen. et sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber		A Nematodini? Macraulacinae false click beetle. The type species is N. balticus.
Palaeocantharis	Gen. et sp. nov	Disputed	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber		A soldier beetle. The type species is P. panna. Fanti & Müller (2022) considered the genus Palaeocantharis to a be a junior synonym of the genus Elektrokleinia, though the authors maintained P. panna as a distinct species within the latter genus.
Podistra guthriei	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber		A soldier beetle.
Poecilochrus balticus	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		A Eucnemini Eucneminae false click beetle.
Potergites brittoni	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		A Dendrocharini Eucneminae false click beetle.
Pseudomataeopsephus^[166]	Gen. et sp. nov	In press	Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber		A water-penny beetle. The type species is P. burmensis.
Pseudopactopus	Gen. et sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber		A throscid false click beetle. The type species is P. robustus.
Ptilodactyla eocenica^[167]	Sp. nov	Valid	Kundrata, Bukejs & Blank in Kundrata et al.	Late Eocene Priabonian	Baltic amber		APtilodactyla species toe winged beetle.
Rhacopus balticus	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		A Rhacopus species eucnemid false click beetle
Rhagomicrus balticus	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		A Dirhagini Melasinae false click beetle.
Rheanischia^[168]	Gen. et sp. nov		Li, Chang & Muona	Early Cretaceous (Hauterivian to Aptian)	Yixian Formation		An Anischiinae false click beetle. The type species is R. brevicornis.
Silis curleri^[169]	Sp. nov	Valid	Fanti & Pankowski	Burdigalian	Dominican amber		A species of Silis.
Silis hegnai	Sp. nov	Valid	Fanti & Pankowski	Burdigalian	Dominican amber		A species of Silis.
Xylophilus balticus	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber		A Xylophilus species eucnemid false click beetle.

=Elateriformia research

=

• A study on the anatomy and phylogenetic relationships of Archaeolus funestus is published by Li et al. (2021).^[170]
• A study on the morphological diversity of larvae of extant and fossil members of the family Brachypsectridae is published by Haug et al. (2021).^[171]

Scarabaeiformia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Goriresina^[172]	Gen. et sp. nov	Valid	Matalin, Perkovsky & Vasilenko	Late Eocene Priabonian	Rovno amber		A tiger beetle. The type species is G. fungifora.
Onthophagus pilauco^[173]	Sp. nov	Valid	Tello et al.	Late Pleistocene	Pilauco Bajo		A scarab beetle.

Staphyliniformia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acritus sutirca^[174]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber		A species of Acritus.
Bacanius gorskii^[175]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber		A species of Bacanius.
Burmagluta^[176]	Gen. et sp. nov	Valid	Yin & Cai	Cretaceous	Burmese amber		A Brachyglutini Pselaphinae rove beetle. The type species is B. rougemonti.
Carinumerus^[177]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber		An Onthophilinae clown beetle. The type species is C. maddisoni.
Charhyphus serratus^[178]	Sp. nov	In press	Yamamoto & Shavrin	Eocene	Baltic amber		A Phloeocharinae rove beetle.
Cretosaja^[179]	Gen. et sp. nov	In press	Sohn & Nam	Early Cretaceous	Jinju Formation		A carrion beetle. The type species is C. jinjuensis.
Druantia	Gen. et sp. nov	Valid	Caterino	Cenomianian	Burmese amber		A Dendrophilinae clown beetle. The type species is D. aeterna.
Euconnus nathani^[180]	Sp. nov	Valid	Jałoszyński & Perkovsky	Eocene	Rovno amber		A Euconnus species Scydmaeninae rove beetle.
Eusphalerum bukejsi^[181]	Sp. nov	Valid	Shavrin in Shavrin & Kairišs	Eocene	Baltic amber		A Eusphalerum species Omaliinae rove beetle.
Megabythinus^[182]	Gen. et sp. nov	Valid	Yin, Zhao & Cai	Cretaceous Cenomanian	Burmese amber		A Bythinini Pselaphinae rove beetle. The type species is M. spinitibialis.
Paroxyporus^[183]	Gen. et sp. nov	Valid	Cai	Early Cretaceous	Yixian Formation		A rove beetle. The type species is P. fossilis.
Phasmister	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber		An Onthophilinae clown beetle. The type species is P. cristatus.
Ponohydrochus^[184]	Gen. et sp. nov	Valid	Prokin & Strelnikova	Early Cretaceous (Hauterivian)			A water scavenger beetle. The type species is P. buryaticus.
Proptomaphaginus alleni^[185]	Sp. nov	Valid	Perreau & Haelewaters in Perreau, Haelewaters & Tafforeau	Miocene Burdigalian	Dominican amber		A Ptomaphagini round fungus beetle.
Trichophya minor^[186]	Sp. nov	Valid	Yamamoto & Newton	Cretaceous (Cenomanian)	Burmese amber		A Trichophya species Trichophyinae rove beetle.
Yethiha	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber		A Dendrophilinae clown beetle. The type species is Y. peregrina.

other Coleopteran research

• A study on the evolution of beetles from the Early Permian to Middle Triassic, evaluating the impact of the Permian–Triassic extinction event on beetle diversity, is published by Zhao et al. (2021).^[187]
• Revision of Notocupes undatabdominus and N. ? multituberatus from the Lower Cretaceous of South China is published by Li, Huang & Cai (2021), who argue that both species do not belong to the genus Notocupes, transfer N. ? multituberatus to the genus Lasiosyne, and consider "N." undatabdominus to be likely a member of the family Artematopodidae.^[188]
• The oldest solid wood-borer larvae reported to date (belonging to the families Buprestidae and possibly Cerambycidae) are described from the Cretaceous amber from Myanmar by Haug et al. (2021).^[189]
• Mitochondrial genomes and nuclear ribosomal DNA of beetles, in some cases identifiable to species, are recovered from pack rat middens from California (United States) and Baja California (Mexico) dating up to ~ 34,355 years before present by Smith et al. (2021).^[190]

Clade Dictyoptera

New Dictyopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Afrophaga^[191]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)			A cockroach belonging to the family Corydiidae. Genus includes new species A. extincta.
Alienopterix mlynskyi^[192]	Sp. nov	In press	Sendi in Vršanský et al.	Cretaceous	Burmese amber		An alienopterid.
Alienopterix smidovae	Sp. nov	In press	Hinkelman in Vršanský et al.	Cretaceous	Burmese amber		An alienopterid.
Arceotermes^[193]	Fam. et gen. et sp. nov	Valid	Engel & Jiang in Jiang et al.	Cretaceous	Burmese amber		A termite belonging to the new family Arceotermitidae. Genus includes new species A. hospitis.
Caligoptera^[194]	Gen. et sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber		A cockroach belonging to the family Ectobiidae. Genus includes new species C. hinkelmani.
Chuanblatta^[195]	Gen. et 2 sp. nov	In press	Liang et al.	Middle Jurassic	Jiulongshan Formation		A cockroach belonging to the family Raphidiomimidae. Genus includes new species C. huntianlingensis and C. lata.
Cuniculoblatta^[196]	Gen. et sp. nov	In press	Hinkelman	Cretaceous	Burmese amber		A cockroach. Genus includes new species C. brevialata .
Ectoovia	Gen. et sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber		A cockroach belonging to the family Ectobiidae. Genus includes new species E. protecta.
? Elisama algeriaensis	Sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)			A member of the family Blattulidae.
Eminespina^[197]	Gen. et sp. nov	Valid	Chen, Zhang Shi	Late Cretaceous (Cenomanian)	Burmese amber		An alienopterid. Genus includes new species E. burma.
Fractalia^[198]	Fam. et gen. et 2 sp. nov	Valid	Hinkelman & Vršanský in Vršanský et al.	Late Jurassic–Late Cretaceous (Kimmeridgian–Cenomanian)	Burmese amber		A member of Corydioidea, the type genus of the new family Fractaliidae Vršanský & Hinkelman. The type species is F. articulata Hinkelman; genus also includes new species F. aristovi Vršanský.
Fragosublatta^[199]	Gen. et sp. nov	Valid	Chen, Shih & Ren in Chen et al.	Late Cretaceous (Cenomanian)	Burmese amber		A cockroach belonging to the family Corydiidae. The type species is F. pectinata.
Glyptotermes abyssinicus^[200]	Sp. nov	Valid	Bouju, Jouault & Perrichot	Early Miocene	Ethiopian amber		A termite belonging to the family Kalotermitidae.
Kinneyblatta^[201]	Gen. et sp. nov	Valid	Schneider et al.	Carboniferous (Kasimovian)	Atrasado Formation		A member of Blattodea belonging to the family Spiloblattinidae. The type species is K. huberi.
Kriedoblatta	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)			A member of the family Liberiblattinidae. Genus includes new species K. gondwanensis.
? Kurablattina samsonovi	Sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)			A member of the family Liberiblattinidae.
Labradormantis^[202]	Gen. et sp. nov	Valid	Demers-Potvin et al.	Late Cretaceous (Cenomanian)	Redmond Formation		A praying mantis belonging to the family Baissomantidae. Genus includes new species L. guilbaulti.
Latiblattella basaltica^[203]	Sp. nov	In press	Vršanský, Poschmann & Vidlička	Late Oligocene			A species of Latiblattella.
Latiblattella karlgruberi	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber
Laticephalana^[204]	Gen. et sp. nov	Valid	Luo et al.	Cretaceous	Burmese amber		A member of the family Umenocoleidae. Genus includes new species L. liuyani.
Lukotermes^[205]	Gen. et sp. nov	Valid	Perkovsky & Nel	Eocene (Priabonian)	Rovno amber		A termite belonging to the family Rhinotermitidae. Genus includes new species L. milescaput.
Meiatermes cretacicus^[206]	Sp. nov	In press	Bezerra et al.	Early Cretaceous	Crato Formation		A termite.
Meloblatta	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)			A member of the family Mesoblattinidae. Genus includes new species M. brezinica.
Milesitermes^[207]	Gen. et sp. nov	Valid	Jouault et al.	Cretaceous	Burmese amber		A termite belonging to the family Mastotermitidae. Genus includes new species M. engeli.
Miniblattina^[208]	Gen. et sp. nov	In press	Sendi	Early Cretaceous (Barremian)	Lebanese amber		A cockroach belonging to the family Liberiblattinidae. Genus includes new species M. libera.
Mongolblatta sendii^[209]	Sp. nov	In press	Hinkelman	Cretaceous	Burmese amber		A member of the family Mesoblattinidae.
Morphna cenozoica^[210]	Sp. nov	In press	Šmídová, Vidlička & Wedmann	Eocene	Messel pit		A cockroach belonging to the family Blaberidae.
Morphna cretacica	Sp. nov	In press	Šmídová, Vidlička & Wedmann	Cretaceous	Burmese amber		A cockroach belonging to the family Blaberidae.
Nadveruzenie	Gen. et sp. nov	In press	Vršanský, Hinkelman & Sendi in Vršanský et al.	Cretaceous	Burmese amber		An alienopterid. Genus includes new species N. postava.
Nodosigalea simplivena^[211]	Sp. nov	In press	Šmídová	Cretaceous	Burmese amber		A cockroach belonging to the family Corydiidae.
Ocelloblattula margarita	Sp. nov	Valid	Koubová & Vršanský in Vršanský et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of Corydioidea.
Olenablatta	Gen. et sp. nov	In press	Šmídová	Cretaceous	Burmese amber		A cockroach belonging to the family Olidae. Genus includes new species O. vrsanskyi.
Otazka	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)			A member of the family Mesoblattinidae. Genus includes new species O. systematicka.
Pabuonqed magna^[212]	Sp. nov	In press	Song et al.	Cretaceous	Burmese amber		A member of Mastotermitoidea belonging to the family Pabuonqedidae.
Paekthoblatta^[213]	Gen. et sp. nov	Valid	So et al.	Early Cretaceous	Sinuiju Formation		A member of the family Raphidiomimidae. Genus includes new species P. coreanica.
Periplaneta (?) perialla^[214]	Sp. nov	Valid	Anisyutkin & Perkovsky	Eocene	Rovno amber		A cockroach, possibly a species of Periplaneta.
Petropterix fukuiensis^[215]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation		A member of the family Umenocoleidae.
Praeblattella arcuata	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation		A member of the family Mesoblattinidae.
Praeblattella inexpecta	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation		A member of the family Mesoblattinidae.
Pseudomantina	Gen. et 2 sp. nov	Valid	Sendi & Vršanský in Vršanský et al.	Late Jurassic–Early Cretaceous (Kimmeridgian–Barremian)	Lebanese amber		A member of the family Liberiblattinidae. The type species is P. occisor Sendi; genus also includes new species P. nigroalba Vršanský.
Reticulitermes grimaldii	Sp. nov	Valid	Jouault et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A termite, a species of Reticulitermes.
Sajda	Gen. et sp. nov	Junior homonym	Vršanský et al.	Cretaceous (Hauterivian?)			A member of Holocompsinae. Genus includes new species S. equatorialis. The generic name is preoccupied by Sajda Dworakowska (1981); Jiang, Xing & Li (2023) coined a replacement name Vrsanskysajda.^[216]
Spinaeblattina baekthoensis^[217]	Sp. nov	Valid	So et al.	Early Cretaceous	Sinuiju Formation		A member of the family Mesoblattinidae.
Stavba delicata	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber		A cockroach belonging to the family Liberiblattinidae.
Stavba magnoculara	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber		A cockroach belonging to the family Liberiblattinidae.
Tanytermitalis	Gen. et sp. nov	Valid	Engel & Cai in Jiang et al.	Cretaceous	Burmese amber		A termite belonging to the family Tanytermitidae. Genus includes new species T. philetaerus.

Dictyopteran research

• A study on the phylogenetic placement of the family Umenocoleidae, based on data from a new specimen of Umenocoleus sinuatus from the Lower Cretaceous Zhonggou Formation (China), is published by Liuo et al. (2021).^[218]
• Description of new alienopterid nymphs from the Cretaceous Burmese amber, and a study on life history and affinities of alienopterids, is published by Luo et al. (2021).^[219]
• A study on the sensory organs of Huablattula hui, and on their implications for the knowledge of the ecology of this cockroach, is published by Taniguchi et al. (2021).^[220]

Clade †Glosselytrodea

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Moltenojurina^[221]	Gen. et sp. nov	Valid	Béthoux & Anderson	Late Triassic (Carnian)	Molteno Formation		A polycytellid Glosselytrodean. The type species is M. parva.
Polycytella rasnitsyni	Sp. nov	Valid	Béthoux & Anderson	Late Triassic (Carnian)	Molteno Formation		A polycytellid Glosselytrodea.

Hymenopterans

New Hymenopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aethemelikertes^[222]	Gen. et sp. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber		A bee belonging to the tribe Melikertini. The type species is A. emunctorii.
Alivespa gracilenta^[223]	Sp. nov	In press	Wu et al.	Cretaceous	Burmese amber		A member of the family Vespidae.
Allommation^[224]	Fam. nov., Gen. et 3 sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Apoidea in the new family Allommationidae. Genus includes new species A. procerum, A. elongatum and A. grande.
Amelikertotes	Gen. et comb. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber Bitterfeld amber		A bee belonging to the tribe Melikertini. The type species is "Melikertes" clypeatus Engel (2001).
Ammosphecium	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Crabroninae. Genus includes new species A. diabolicum.
Anochetus miosumatrensis^[225]	Sp. nov	Valid	Ngô-Muller, Garrouste & Nel in Ngô-Muller et al.	Miocene			An ant, a species of Anochetus.
Archaeovespa incompleta	Sp. nov	In press	Wu et al.	Cretaceous	Burmese amber		A member of the family Vespidae.
Archeonesia^[226]	Gen. et sp. nov	Valid	Tribull, Pankowski & Colombo	Eocene	Baltic amber		A Pristocerinae Bethylid. The type species is A. eocena.
Arkadiserphus^[227]	Gen. et sp. nov	Valid	Rasnitsyn	Late Jurassic	Karabastau Formation		A member of Proctotrupoidea belonging to the family Peleserphidae. The type species is A. leleji.
Ascogaster (Syntaphus) latitibialis^[228]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Cheloninae.
Aulacicosmus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Heterogynaidae. Genus includes new species A. setosus.
Austroplebeia fujianica^[229]	Sp. nov	Valid	Engel in Engel et al.	Miocene (Langhian)	Zhangpu amber		A stingless bee, a species of Austroplebeia.
Burmabracon^[230]	Gen. et 2 sp. nov	Valid	Li et al.	Cretaceous	Burmese amber		A member of the family Braconidae belonging to the subfamily Protorhyssalinae. Genus includes new species B. gracilens and B. grossus.
Burmapyris^[231]	Gen. et sp. nov	Valid	Jouault, Perrichot & Nel	Cretaceous			A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species B. azevedoi.
Burmoxyela^[232]	Gen. et sp. nov	Valid	Zheng et al.	Late Cretaceous (Cenomanian)	Burmese amber		A sawfly belonging to the family Syspastoxyelidae. Genus includes new species B. lii.
Burmusculus angustus^[233]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber		A member of Apocrita belonging to the family Burmusculidae.
Buserphites^[234]	Gen. et 2 sp. nov	In press	Herbert & McKellar	Cretaceous	Burmese amber		A member of the family Serphitidae. Genus includes new species B. applanatus and B. myanmarensis.
Chainochora^[235]	Gen. et sp. nov	Valid	Chen, van Achterberg & Hong	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is C. syntoma.
Chlorepyris deploegi	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber		A member of the family Bethylidae.
Chlorepyris engeli	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae.
Chlorepyris hopei	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae.
Chlorepyris jouaulti	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber		A member of the family Bethylidae.
Chlorepyris mckellari	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber		A member of the family Bethylidae.
Chlorepyris meunieri	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae.
Chlorepyris neli	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber Rovno amber		A member of the family Bethylidae.
Chlorepyris perrichoti	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae.
Christophus^[236]	Gen. et 2 sp. nov	In press	Brazidec et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Diapriidae. Genus includes new species C. myanmarensis and C. ohmkuhnlei.
Cretastatus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Astatinae. Genus includes new species C. longevus.
Cretorussus^[237]	Gen. et sp. nov	In press	Jouault, Perrichot & Nel	Cretaceous	Burmese amber		A member of Orussoidea. Genus includes new species C. vilhelmseni.
Curiosivespa sennlaubi^[238]	Sp. nov	Valid	Jouault, Rasnitsyn & Nel in Jouault et al.	Early Cretaceous	Crato Formation		A member of the family Vespidae.
Diameneura^[239]	Gen. et sp. nov	In press	Santer & Álvarez-Parra in Santer et al.	Early Cretaceous (Albian)	Escucha Formation		A member of the family Spathiopterygidae. The type species is D. marveni.
Drymomyrmex rasnitsyni^[240]	Sp. nov	Valid	Radchenko	Late Eocene	Rovno amber		An ant belonging to the subfamily Formicinae.
Elasmosomites arkadyleleji^[241]	Sp. nov	Valid	Belokobylskij in Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Euphorinae.
Electrofoenops exaltatus^[242]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber		A member of the family Aulacidae.
Embolemus brachypterus^[243]	Sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber		A member of the family Embolemidae.
Embolemus micropterus	Sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber		A member of the family Embolemidae.
Eopelecinus marechali^[244]	Sp. nov	Valid	Jouault	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Pelecinidae.
Eoteredon^[245]	Gen. et sp. nov	Valid	Archibald, Aase & Nel	Early Eocene	Green River Formation		A horntail. Genus includes new species E. lacoi.
Ephedrus rasnitsyni^[246]	Sp. nov	Valid	Davidian & Kaliuzhna in Davidian, Kaliuzhna & Perkovsky	Middle Eocene	Sakhalinian amber		A member of the family Braconidae belonging to the subfamily Aphidiinae.
Exilaulacus advenus	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber		A member of the family Aulacidae.
Exilaulacus eximius	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber		A member of the family Aulacidae.
Formica ribbeckei^[247]	Sp. nov	Valid	Radchenko & Perkovsky in Radchenko, Perkovsky & Vasilenko	Late Eocene	Rovno amber		An ant, a species of Formica.
Glenocephalus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Apoidea in the new family Cirrosphecidae. Genus includes new species G. mandibularis.
Gloxinius^[248]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae; a new genus for "Epyris" bifossatus Brues (1939).
Grandixyela^[249]	Gen. et sp. nov	Valid	Zheng in Zheng et al.	Cretaceous	Burmese amber		A member of the family Syspastoxyelidae. The type species is G. rasnitsyni.
Haidomelikertes	Gen. et sp. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber		A bee belonging to the tribe Melikertini. The type species is H. uraeus.
Haptodioctes	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Apoidea in the new family Cirrosphecidae. Genus includes new species H. apiformis.
Helosericus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Psenini. Genus includes new species H. verrucosus.
Heteroichneumon^[250]	Gen. et sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber		A member of the family Ichneumonidae. The type species is H. rasnitsyni.
Heteropimpla pulverulenta	Sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber		A member of the family Ichneumonidae.
Holepyris rasnitsyni	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae.
Holepyris terayamai	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber		A member of the family Bethylidae.
Holopsenella antiqua^[251]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber		A member of Aculeata of uncertain phylogenetic placement.
Holopsenella burmitica	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber		A member of Aculeata of uncertain phylogenetic placement.
Holopsenella gothica	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber		A member of Aculeata of uncertain phylogenetic placement.
Hybristodryinus castaneus^[252]	Sp. nov	Valid	Olmi, Guglielmino & Chen in Olmi et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Dryinidae.
Hybristodryinus viriosus^[253]	Sp. nov	Valid	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Dryinidae.
Hybristodryinus zaifui	Sp. nov	Valid	Chen, Olmi & Perkovsky in Olmi et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Dryinidae.
Janzenella theia^[254]	Sp. nov	Valid	Bremer & Talamas in Bremer, van de Kamp & Talamas	Eocene	Baltic amber		A species of Janzenella.
Karataus ryonsangensis^[255]	Sp. nov	In press	Won & So	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation		A wasp belonging to the family Ephialtitidae.
Keratodellitha^[256]	Gen. et 3 sp. nov	Valid	Jouault et al.	Cretaceous	Burmese amber		A member of Evanioidea belonging to the family Othniodellithidae. Genus includes new species K. anubis, K. basilisci and K. kirin.
Kleistochora	Gen. et sp. nov	Valid	Chen, van Achterberg & Hong	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is K. dolichura.
Lelejus^[257]	Gen. et sp. nov	Valid	Radchenko & Proshchalykin	Late Eocene	Rovno amber		An ant belonging to the subfamily Myrmicinae. The type species is L. venustus.
Lonchodryinus groehni^[258]	Sp. nov	Valid	Olmi et al.	Late Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Dryinidae.
Mael	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber		A member of the family Bethylidae belonging to the subfamily Scleroderminae; a new genus for "Calyoza" longiceps Brues (1923).
Megalyrhyssalus^[259]	Gen. et sp. nov	Valid	Belokobylskij & Jouault	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Braconidae. Genus includes new species M. clavicornis.
Megarolium	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Bembicinae. Genus includes new species M. paradoxum.
Merascylla	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Bethylidae belonging to the subfamily Pristocerinae; a new genus for "Epyris" atavellus Cockerell (1920).
Mesoserphites	Gen. et 5 sp. nov	In press	Herbert & McKellar	Cretaceous	Burmese amber		A member of the family Serphitidae. Genus includes new species M. annulus, M. giganteus, M. engeli, M. scutatus and M. viraneacapitis.
Meteorus arasnitsyni	Sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Meteorus.
Microtypus eocenus	Sp. nov	Valid	Belokobylskij et al.	Eocene	Rovno amber		A member of the family Braconidae belonging to the subfamily Microtypinae.
Mintara	Gen. et sp. nov	In press	Brazidec et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Diapriidae. Genus includes new species M. parva.
Odontosericus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Psenini. Genus includes new species O. burmensis.
Orthosyntexis^[260]	Gen. et 2 sp. nov	Valid	Gao, Engel, Shih, Gao in Gao et al.	Cretaceous	Burmese amber		A member of the family Anaxyelidae. The type species is O. elegans; genus also includes O. thanti.
Palaeocolastes	Gen. et sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Exothecinae. The type species is P. bruesi.
Paleoaulacus^[261]	Gen. et sp. nov	Valid	Jouault & Nel	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Aulacidae. Genus includes new species P. minutus.
Paleoxyela^[262]	Gen. et sp. nov	Valid	Jouault, Aase & Nel	Eocene (Chadronian)	Florissant Formation		A member of the family Xyelidae. The type species is P. nearctica.
Peleserphus qufuensis^[263]	Sp. nov	In press	Yu et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Peleserphidae.
Pheidole pauchil^[264]	Sp. nov	Valid	Varela-Hernández & Riquelme	Miocene	Mexican amber		An ant, a species of Pheidole.
Pinguixyela	Gen. et comb. nov	Valid	Zheng et al.	Late Cretaceous (Cenomanian)	Burmese amber		A sawfly belonging to the family Syspastoxyelidae; a new genus for "Syspastoxyela" pinguis.
Platyxyela tenuis^[265]	Sp. nov	Valid	Zheng et al.	Middle Jurassic (Callovian-Bathonian boundary)	Jiulongshan Formation		A member of the family Xyelidae.
Pompilopterus ryonsangensis	Sp. nov	In press	Won & So	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation		A wasp belonging to the family Angarosphecidae.
Pristinopterus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Bembicinae. Genus includes new species P. spectabilis.
Pristopsen	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Psenini. Genus includes new species P. obscurus.
Prosyntexis sennlaubi^[266]	Sp. nov	Valid	Jouault & Nel	Early Cretaceous (Aptian)	Crato Formation		A member of the family Sepulcidae.
Protopyris	Gen. et sp. nov	Valid	Jouault & Nel in Jouault, Perrichot & Nel	Early Cretaceous (Albian)	Hkamti amber		A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species P. myanmarensis.
Prototeleia^[267]	Gen. et sp. nov	Valid	Talamas et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Platygastridae. The type species is P. kleio.
Proxyelia	Gen. et sp. nov	Valid	Jouault, Aase & Nel	Eocene (Ypresian)	Green River Formation		A member of the family Xyelidae. The type species is P. pankowskii.
Ptilocosmus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Heterogynaidae. Genus includes new species P. corniculatus.
Ramageoptera^[268]	Gen. et sp. nov	Valid	Jouault & Brazidec	Cretaceous	Burmese amber		A member of the family Bethylidae belonging to the subfamily Protopristocerinae. Genus includes new species R. platycephala.
Rasnichneumon	Gen. et 2 sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber		A member of the family Ichneumonidae. The type species is R. alexandri; genus also includes R. gracilis.
Rasnitsynum^[269]	Gen. et sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber		A member of the family Dryinidae. Genus includes new species R. burmense.
Rogichneumon	Gen. et sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber		A member of the family Ichneumonidae. The type species is R. braconidicus.
Rovenosa alexrasnitsyni^[270]	Sp. nov	Valid	Manukyan in Manukyan & Zhindarev	Late Eocene	Prussian Formation (Baltic amber)		A member of the family Ichneumonidae.
Rovenosa khalaimi	Sp. nov	Valid	Manukyan in Manukyan & Zhindarev	Late Eocene	Prussian Formation (Baltic amber)		A member of the family Ichneumonidae.
Sakhalinencyrtus^[271]	Gen. et sp. nov	Valid	Simutnik in Simutnik, Perkovsky & Vasilenko	Middle Eocene	Sakhalinian amber		A member of the family Encyrtidae. The type species is S. leleji.
Sakhalinoctonus^[272]	Gen. et sp. nov	Valid	Davidian in Davidian, Manukyan & Belokobylskij	Middle Eocene	Sakhalinian amber		A member of the family Braconidae belonging to the subfamily Aphidiinae. Genus includes new species S. alexrasnitsyni.
Scleroxyela	Gen. et sp. nov	Valid	Zheng et al.	Middle Jurassic (Callovian-Bathonian boundary)	Jiulongshan Formation		A member of the family Xyelidae. The type species is S. daohugouensis.
Spalangiopelta darlingi^[273]	Sp. nov	Valid	Moser in Moser et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Pteromalidae.
Spalangiopelta semialba	Sp. nov	Valid	Moser in Moser et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Pteromalidae.
Spheciellus	Fam. nov., Gen. et sp. nov.	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of Apoidea in the new family Spheciellidae. Genus includes new species S. aenigmaticus.
Stephanorhyssalus	Gen. et sp. nov	Valid	Belokobylskij & Jouault	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Braconidae. Genus includes new species S. longiscapus.
Tetragonula florilega	Sp. nov	Valid	Engel in Engel et al.	Miocene (Langhian)	Zhangpu amber		A stingless bee, a species of Tetragonula.
Thigmocosmus	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Heterogynaidae. Genus includes new species T. longicornis.
Tyrannomecia^[274]	Gen. et sp. nov	In press	Jouault & Nel	Paleocene	Menat Formation		An ant belonging to the stem group of the subfamily Myrmeciinae. Genus includes new species T. inopinata.
Vespula? hassiaca^[275]	Sp. nov	In press	Abels & Wedmann	Eocene	Messel pit		A member of the family Vespidae.
Xeris dorbnikensis^[276]	Sp. nov	Valid	Manukyan in Manukyan & Smirnova	Eocene	Baltic amber	Europe (Baltic Sea region)	A horntail. Originally described as a species of Xeris; Archibald & Rasnitsyn (2022) transferred it to the genus Urocerus.^[277]
Zigrasimecia goldingot^[278]	Sp. nov	In press	Zhuang et al.	Cretaceous	Burmese amber		An ant in the tribe Sphecomyrmini.

Hymenopteran Research

• Guo et al. (2021) report the discovery of two queen ant specimens from Burmese amber, belonging to the species Haidomyrmex cerberus and including the first known alate queen in Haidomyrmex.^[279]
• Cooccurrence of ants belonging to extant Holarctic genus Lasius and a tropical weaver ant in the same piece of amber is reported from the Bitterfeld amber (Germany) by Radchenko & Perkovsky (2021).^[280]
• Original claims that both Melittosphex burmensis and Discoscapa apicula were primitive bees have been rejected by Rosa & Melo, who instead consider the former to be an aculeate wasp of uncertain affinity, and the latter to belong within the crabronid wasp subfamily Crabroninae, a lineage not directly related to bees.

Clade Neuropterida

Neuropterans

New Neuropteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Ankyloleon^[281]	Gen. et sp. nov	Valid	Badano, Haug & Cerretti in Badano et al.	Late Cretaceous (Cenomanian)	Burmese amber		An early representative of the clade Myrmeleontiformia. The type species is A. caudatus.
Astioberotha coutreti^[282]	Sp. nov	In press	Jouault	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Rhachiberothidae belonging to the subfamily Paraberothinae.
Burmotachinymphes^[283]	Gen. et sp. nov	Valid	Cao, Wang & Liu	Cretaceous	Burmese amber		A member of the family Mesochrysopidae. Genus includes new species M. bilobata.
Calobabinskaia^[284]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Babinskaiidae. The type species is C. xiai.
Fuscopolystoechotes^[285]	Gen. et sp. nov	Valid	Xu, Zheng, Shih & Wang in Xu et al.	Middle Jurassic	Jiulongshan Formation		A member of the family Ithonidae. Genus includes new species F. reticulatus.
Karenina cuneiformis^[286]	Sp. nov	Valid	Nakamine	Early Cretaceous (Aptian)	Crato Formation		A member of the family Mesochrysopidae.
Lichenipolystoechotes fenestratus	Sp. nov	Valid	Xu, Fang, Shih & Wang in Xu et al.	Middle Jurassic	Jiulongshan Formation		A member of the family Ithonidae.
Osmylidia donnae^[287]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Quilchena site
Osmylidia glastrai	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Klondike Mountain Formation
Osmylidia picta	Comb. nov	Valid	(Hagen)	Priabonian	Baltic amber		A protosmyline Osmylid lance lacewing Moved from "Protosmylus" picta (1913)
Osmylidia taliae	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Green River Formation Parachute Creek Member
Osmyloberotha^[288]	Gen. et sp. nov	Valid	Khramov	Late Cretaceous (Cenomanian)	Burmese amber		A beaded lacewing. Genus includes new species O. simpla.
Paradoxoberotha^[289]	Gen. et sp. nov	In press	Nakamine et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Rhachiberothidae. Genus includes new species P. chimaera.
Paraneliana^[290]	Gen. et sp. nov	Valid	Jouault & Nel	Early Cretaceous	Crato Formation		A member of the family Babinskaiidae. Genus includes new species P. sennlaubi.
Rasnitsyneura^[291]	Gen. et comb. nov	Valid	Makarkin, Ansorge & Khramov	Early Jurassic (Toarcian)			A member of the family Ithonidae sensu lato belonging to the subfamily Epigambriinae. The type species is "Prohemerobius" aequabilis Bode (1953).
Sinonothochrysa^[292]	Gen. et sp. nov	In press	Huang & Nel in Huang et al.	Paleocene	Buxin Formation		A member of the family Chrysopidae. Genus includes new species S. zhangi
Stenobabinskaia	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Babinskaiidae. The type species is S. punctata.
Xiaobabinskaia	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Babinskaiidae. The type species is X. lepidotricha.

Neuropteran research

• Review of the known record of spoon-winged lacewing larvae is published Haug, Haug & Haug (2021), who report two specimens from the Cretaceous amber from Myanmar which might represent the first known larvae of members of the subfamily Nemopterinae in the fossil record.^[293]
• New lacewing larvae with elongated mouthparts are described from the Cretaceous amber from Myanmar by Haug et al. (2021), who also review the morphological diversity of fossil lacewing larvae.^[294]

Raphidiopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Archiinocellia protomaculata^[295]	comb nov	valid	(Engel)	Lutetian	Green River Formation		A raphidiid snakefly. Moved from Archiinocellia protomaculata Engel (2011)^[296]
Electrobaissoptera liui^[297]	Sp. nov	In press	Jouault, Engel & Nel	Late Cretaceous (Cenomanian)	Burmese amber		A baissopterid snakefly.
Megaraphidia antiquissima	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Tranquille Formation		A raphidiid snakefly.
Megaraphidia hopkinsi	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Allenby Formation		A raphidiid snakefly.
Megaraphidia klondika	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Klondike Mountain Formation		A raphidiid snakefly.
Megaraphidia ootsa	Sp. nov	Valid	Archibald & Makarkin	Ypresian	"Unnamed unit" Ootsa Group		A raphidiid snakefly.
Mesoraphidia baekthoensis^[298]	Sp. nov	In press	Ri et al.	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation		A member of the family Mesoraphidiidae.
Mesoraphidia ryonsangensis	Sp. nov	In press	Ri et al.	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation		A member of the family Mesoraphidiidae.

Clade †Palaeodictyopteroidea

†Palaeodictyoptera

New Palaeodictyopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Dictyoneura goujonorum^[300]	Sp. nov	In press	Logghe et al.	Carboniferous-Permian (Gzhelian-Asselian)			A member of Palaeodictyoptera.
Sandiella herbigi^[301]	Sp. nov	In press	Brauckmann, Herd & Leipner	Carboniferous (Moscovian)			A member of Palaeodictyoptera belonging to the family Eugereonidae.

Palaeodictyopteran research

• A study on the anatomy of fossil specimens described as consecutive series of different larval stages of palaeodictyopteran species Tchirkovaea guttata and Paimbia fenestrata, evaluating their implications for the knowledge of wing development in palaeodictyopterans, is published by Rosová, Sinitshenkova & Prokop (2021).^[302]

Clade Palaeoptera

Ephemeroptera

New Ephemeropteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Astraeoptera^[303]	Gen. et sp. nov	Valid	Brandão et al.	Early Cretaceous	Crato Formation		A mayfly of uncertain phylogenetic placement. Genus includes new species A. cretacica.
Burmella^[304]	Gen. et 2 sp. nov	Valid	Godunko, Martynov & Staniczek	Late Cretaceous (Cenomanian)	Burmese amber		A vietnamellid mayfly. The type species is B. paucivenosa; genus also includes B. clypeata.
Protoligoneuria heloisae^[305]	Sp. nov	In press	Storari et al.	Early Cretaceous (Aptian)	Crato Formation		A hexagenitid mayfly.
Triassodotes rasnitsyni^[306]	Sp. nov	Valid	Sinitshenkova	Triassic			A misthodotid mayfly.

Ephemeropteran research

• Sroka et al. (2021) provide new information on the morphology of Permian stem-mayflies Misthodotes sharovi and M. zalesskyi from the Tshekarda locality (Kungurian Koshelevka Formation; Perm Krai, Russia), and infer the life history traits of both the adult and larval stages of these insects.^[307]
• Possible underwater mayfly burrows are described from the Turonian Ferron Sandstone (Utah, United States) by King et al. (2021), who interpret this finding as evidence that burrowing mayflies adapted to domicile filter-feeding during or prior to the Turonian, and name a new ichnospecies Glossifungites gingrasi.^[308]

Odonatoptera

Odonata

New Odonatan taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Chalcolestes tibetensis^[309]	Sp. nov	In press	Xia et al.	Late Eocene	Nima Basin		A species of Chalcolestes.
Cretaphylolestes^[310]	Gen. et sp. nov	In press	Huang et al.	Early Cretaceous (Aptian)	Shouchang Formation		A damselfly belonging to the family Synlestidae. Genus includes new species C. cretacicus.
Dakotaeschnidium^[311]	Gen. et sp. nov	Valid	Nel	Late Cretaceous (Maastrichtian)	Fox Hills Formation		A dragonfly belonging to the family Aeschnidiidae. Genus includes new species D. spedeni.
Dysagrion pruettae^[312]	Sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation		A member of the family Dysagrionidae.
Dysagrionites	Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation		A collective group name for any fossil species referable to the Dysagrioninae whose orthotaxon generic affinity is unclear. Includes new species D. delinei.
Epilestes rasnitsyni^[313]	Sp. nov	Valid	Felker & Vasilenko	Permian			A damselfly belonging to the family Permagrionidae.
Jingguaeshna^[314]	Gen. et sp. nov	Valid	Zheng et al.	Miocene			A dragonfly belonging to the group Aeshnoidea, possibly a member of the family Telephlebiidae. Genus includes new species J. taoae.
Marado^[315] ^[316]	Gen. et sp. nov	Valid	Petrulevičius	Paleocene	Maíz Gordo Formation		A damselfly belonging to the family Coenagrionidae. The type species is M. marado.
Moltenophlebia^[317]	Gen. et sp. nov	Valid	Deregnaucourt et al.	Triassic	Molteno Formation		A member of the stem group of Odonata, belonging to the group Zygophlebiida within the Triadophlebiomorpha. Genus includes new species M. lindae.
Okanagrion	Gen. et 8 sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation		A member of the family Dysagrionidae. The type species is O. threadgillae; genus also includes new species O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum and O. worleyae.
Okanopteryx	Gen. et 3 sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Tranquille Formation		A member of the family Dysagrionidae. The type species is O. macabeensis; genus also includes new species O. fraseri and O. jeppesenorum.
Phyonganpodagrion^[318]	Gen. et sp. nov	In press	So & Won	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation		A damselfly in the family Megapodagrionidae. Genus includes new species P. ryonsangae.
Protohemiphlebia^[319]	Fam. et gen. et 2 sp. nov	In press	Zheng, Jarzembowski & Nel in Zheng et al.	Cretaceous	Burmese amber		A stem hemiphlebioid damselfly belonging to the new family Protohemiphlebiidae. Genus includes new species P. zhangi and P. meiyingae.
Pulchrairina^[320]	Gen. et sp. nov	In press	Martynov, Vasilenko & Perkovsky	Late Eocene	Rovno amber		A damselfly belonging to the group Coenagrionoidea. Genus includes new species P. electra.
Republica^[321]	Gen. et sp. nov	Valid	Archibald & Cannings	Eocene (Ypresian)	Klondike Mountain Formation		A euphaeid damselfly. Type species is R. weatbrooki. Not to be confused with the plant genus Republica also from the formation.	thumb|center|upright|Republica weatbrooki
Stenodiafanus	Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation		A dysagrionid odonate. Type species is S. westersidei.
Stenophlebia ryonsangensis^[322]	Sp. nov	In press	Won, So & Kim	Early Cretaceous	Sinuiju Formation		A stenophlebiid odonate.
Whetwhetaksa	Fam. Nov., Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation		The type genus of Whetwhetaksidae, a new family of uncertain placement. The type species is W. millerae.

Odonate research

• Archibald et al. (2021) establish a new, extinct suborder of Odonata named Cephalozygoptera, close to but distinct from damselflies, composed of the families Dysagrionidae, Sieblosiidae and possibly Whetwhetaksidae; their study is subsequently criticized by Nel & Zheng (2021), who consider evidence presented by Archibald et al. to be insufficient to support a new clade, and reject the suborder Cephalozygoptera as unfounded.^{[323] [324]}

†Protozygoptera

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Kennedya kedrovkensis^[325]	Sp. nov	Valid	Felker	Early Triassic			A member of Protozygoptera belonging to the family Kennedyidae.
Kennedya suchonensis^[326]	Sp. nov	Valid	Felker	Late Permian
Progoneura kemerovensis	Sp. nov	Valid	Felker	Early Triassic			A member of Protozygoptera belonging to the family Kennedyidae.

Other odonatopterans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Brunellopteron^[327]	Gen. et sp. nov	Valid	Béthoux, Deregnaucourt & Norrad in Béthoux et al.	Carboniferous (Moscovian)	Sunbury Creek Formation		A member of Neodonatoptera of uncertain phylogenetic placement. The type species is B. norradi.
Osnabruggiala^[328]	Gen. et sp. nov	Valid	Zessin, Brauckmann & Gröning	Carboniferous (Pennsylvanian)	Osnabrück Formation		Probably a basal member of Odonatoptera. The type species is O. seppelti.

Clade †Paoliidea

†Paoliida

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Avionblattinopsis^[329]	Gen. et sp. nov	In press	Quispe et al.	Carboniferous (Moscovian)			A blattinopsid Paoliidan. The type species is A. oudardi.
Blattinopsis indefinitus^[330]	Sp. nov	Valid	Rasnitsyn & Aristov	Permian (Kungurian)			A Blattinopsidae Paoliidan
Glaphyrophlebia borea^[331]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation		A Blattinopsid.
Glaphyrophlebia iva^[332]	Sp. nov	Valid	Aristov & Rasnitsyn	Middle Permian			A member of the family Blattinopsidae.
Glaphyrophlebia komia	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation		A member of the family Blattinopsidae.
Glaphyrophlebia vorkutensis	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation		A member of the family Blattinopsidae.

Clade Paraneoptera

Hemipterans

New Hemipteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acrotiara^[333]	Trib. et gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Cixiidae; the type genus of the new tribe Acrotiarini. The type species is A. multigranulata Luo & Bourgoin.
Aneurus damzeni^[334]	Sp. nov	Valid	Heiss in Heiss, Kairišs & Bukejs	Eocene	Baltic amber		A species of Aneurus.
Aphelicophontes^[335]	Gen. et sp. nov	Valid	Swanson et al.	Eocene	Green River Formation		A member of the family Reduviidae belonging to the subfamily Harpactorinae. Genus includes new species A. danjuddi.
Baltiomiris^[336]	Gen. et sp. nov	Valid	Kim, Chérot & Jung	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Miridae. The type species is B. herczeki.
Barremixius	Gen. et comb. nov	In press	Bourgoin & Szwedo in Luo et al.	Early Cretaceous	Weald Clay		A member of the family Cixiidae; a new genus for "Cixius" petrinus Fennah.
Beaconiella cincta^[337]	Sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation		A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.
Beaconiella pulchra	Sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation		A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.
Burmacader bicoloripennis^[338]	Sp. nov	Valid	Souma, Yamamoto & Takahashi	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Tingidae.
Burmacader grandis^[339]	Sp. nov	Valid	Heiss & Guilbert	Cretaceous	Burmese amber		A member of the family Tingidae.
Burmala^[340]	Gen. et sp. nov	Valid	Liu et al.	Cretaceous	Burmese amber		A member of the family Malmopsyllidae. Genus includes new species B. liaoyaoi.
Burmaprosbole^[341]	Gen. et sp. nov	Valid	Qiao et al.	Cretaceous	Burmese amber		A member of the family Tettigarctidae. Genus includes new species B. nervosa.
Burmavianaida^[342]	Gen. et sp. nov	Valid	Souma, Yamamoto & Takahashi	Cretaceous (Albian or Cenomanian)	Burmese amber		A member of the family Tingidae. The type species is B. anomalocapitata.
Cretamystilus^[343]	Gen. et sp. nov	Valid	Kim, Lim & Jung	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Miridae belonging to the tribe Mecistoscelini. Genus includes new species C. herczeki.
Cretodorus rostellatus^[344]	Sp. nov	In press	Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Mimarachnidae.
Cretomultinervis^[345]	Gen. et sp. nov	Valid	Fu & Huang	Cretaceous	Burmese amber		A member of the family Sinoalidae. Genus includes new species C. burmensis.
Dachibangus hui^[346]	Sp. nov	Valid	Zhang, Yao & Pang in Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Mimarachnidae. Originally described as a species of Dachibangus, but subsequently transferred to the genus Xiaochibangus.^[347]
Delphitiara	Gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is D. tibiocoronata Luo & Bourgoin.
Dorytocus jiaxiaoae^[348]	Sp. nov	Valid	Song, Szwedo & Bourgoin in Song et al.	Late Cretaceous (Cenomanian)	Burmese amber		A planthopper.
Duyana^[349]	Gen. et sp. nov	Valid	Chen et al.	Late Cretaceous (Cenomanian)	Burmese amber		A leafhopper belonging to the subfamily Ledrinae. Genus includes new species D. electronica.
Electromyiomma herczeki^[350]	Sp. nov	Valid	Kim & Jung	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Miridae.
Eospinosus^[351]	Gen. et 2 sp. nov	Valid	Wedmann et al.	Eocene	Messel pit		A member of the family Pentatomidae. The type species is E. peterkulkai; genus also includes E. greenriverensis.
Ferriantenna^[352]	Gen. et sp. nov	Valid	Cumming & Le Tirant	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Coreidae belonging to the subfamily Coreinae. The type species is F. excalibur.
Hallex martinsnetoi^[353]	Sp. nov	In press	Monferran et al.	Early Cretaceous	Crato Formation		A leafhopper.
Hypsotsijilia^[354]	Gen. et sp. nov	Valid	Cifuentes-Ruiz & Brailovsky in Cifuentes-Ruiz et al.	Early Miocene	La Quinta Formation (Mexican amber)		A member of the family Schizopteridae. The type species is H. bretoni.
Inoderbe^[355]	Fam. et gen. et sp. nov	Valid	Shcherbakov & Emeljanov	Cretaceous (Albian–Cenomanian)	Burmese amber		A planthopper belonging to the new family Inoderbidae. The type species is I. rapunzel.
Juroala pulchra^[356]	Sp. nov	Valid	Fu & Huang	Late Jurassic	Daohugou Beds		A member of the family Sinoalidae.
Karatauella^[357]	Nom. nov	Valid	Doweld	Jurassic (Bathonian-Oxfordian)			A member of the family Protopsyllidiidae; a replacement name for Carpenterella Becker-Migdisova (1968).
Latidorsum^[358]	Gen. et sp. nov		Wang, Tang & Yao in Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Tingidae. The type species is L. carinbifarium.
Microtingis^[359]	Gen. et sp. nov	Junior synonym	Heiss & Golub	Cretaceous	Burmese amber		A member of the family Tingidae. Genus includes new species M. leptosoma. Subsequently Heiss & Golub (2021) considered this species to be a junior synonym of Burmavianaida anomalocapitata.^[360]
Multistria	Gen. et sp. nov	Valid	Zhang, Yao & Pang in Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Mimarachnidae. The type species is M. orthotropa.
Paleocader balticus^[361]	Sp. nov	Valid	Heiss & Golub	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Tingidae.
Paleocader pulchellus	Sp. nov	Valid	Heiss & Golub	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Tingidae.
Paleocader rovnensis	Sp. nov	Valid	Heiss & Golub	Eocene	Rovno amber		A member of the family Tingidae.
Parasinalda sukachevae^[362]	Sp. nov	Valid	Golub, Perkovsky & Vasilenko	Late Eocene	Rovno amber		A member of the family Tingidae.
Pentacarinus	Gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is P. kachinensis Luo & Bourgoin.
Pseudocaulisoculus^[363]	Gen. et sp. nov	Valid	Kóbor & Roca-Cusachs	Cretaceous	Burmese amber		A member of the family Yuripopovinidae. Genus includes new species P. longicornis.
Pubivetanthocoris^[364]	Gen. et sp. nov	In press	Tang, Wang & Yao in Tang et al.	Cretaceous	Burmese amber		A member of the family Vetanthocoridae. Genus includes new species P. carinalis.
"Pyrrhocoris" rottensis^[365]	Nom. nov	Valid	Kment	Late Oligocene			A member of Lygaeoidea of uncertain phylogenetic placement; a replacement name for Pyrrhocoris tibialis Statz & Wagner (1950).
Rovnoxestus^[366]	Gen. et sp. nov	Valid	Dietrich, Dmitriev & Perkovsky	Late Eocene	Rovno amber		A leafhopper. Genus includes new species R. rasnitsyni.
Saltissus fennahi^[367]	Sp. nov	In press	Luo, Liu & Jarzembowski in Luo et al.	Early Cretaceous	Weald Clay		A member of the family Mimarachnidae.
Scytinoptera tongchuanensis^[368]	Sp. nov	In press	Zhang, Zheng & Zhang in Zhang et al.	Middle Triassic	Tongchuan Formation		A member of the family Scytinopteridae.
Sinogranulus^[369]	Gen. et sp. nov	In press	Zhang, Chen & Zhang	Middle Triassic	Tongchuan Formation		A member of Scytinopteroidea belonging to the family Granulidae. Genus includes new species S. qishuiheensis.
Spinonympha^[370]	Gen. et sp. nov	Valid	Luo, Wang & Jarzembowski	Late Cretaceous (Cenomanian)	Burmese amber		A planthopper of uncertain phylogenetic placement. The type species is S. shcherbakovi.
Szeiinia^[371]	Fam. nov., Gen. et sp. nov	Valid	Zhang et al.	Late Triassic	Yanchang Formation		A planthopper in the new family Szeiiniidae. Genus includes new species S. huanglongensis
Tennentsia orientalis^[372]	Sp. nov	In press	Fu, Azar & Huang	Middle Triassic	Yanchang Formation		A member of Cicadomorpha belonging to the family Dysmorphoptilidae.
Triknightia	Gen. et sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation		A member of Cicadomorpha belonging to the superfamily Scytinopteroidea and the family Paraknightiidae. The type species is T. mira.
Tuthillia maculosa^[373]	Sp. nov	Valid	Poinar & Brown		Dominican amber		A member of Psylloidea.
Valdicopis^[374]	Gen. et sp. nov	Valid	Li, Chen & Jarzembowski	Early Cretaceous			A member of the family Procercopidae. Genus includes new species V. tonyi.
Youngeewa simillima^[375]	Sp. nov	Valid	Gębicki & Walczak in Gębicki, Walczak & Zmarzły	Eocene	Baltic amber	Europe (Baltic Sea region)	A leafhopper belonging to the subfamily Mileewinae.

Hemipteran research

• Grimaldi & Vea (2021) reinterpret Mesophthirus engeli as a scale insect, and reject the interpretation of this insect as an ectoparasite.^[376]
• Redescription of the holotype specimen of Paranoika placida is published by Moura-Júnior et al. (2021), who transfer this species to the genus Lethocerus, and consider this species to be the senior synonym of Lethocerus vetus.^[377]
• A fulgorid specimen, representing the oldest occurrence of this family in the fossil record in Asia, is described from the Eocene Lunpola Basin (Qinghai-Tibetan Plateau, China) by Xu et al. (2021).^[378]
• Revision of the fossil record of Fulgoromorpha and Cicadomorpha from the Jurassic and Cretaceous of eastern Asia is published by Chen (2021).^[379]

Psocodeans

New Psocodean taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Burmesopsocus^[380]	Gen. et sp. nov	Valid	Yoshizawa in Yoshizawa & Yamamoto	Cretaceous (Albian–Cenomanian)	Burmese amber		A member of Homilopsocidea of uncertain phylogenetic placement. The type species is B. lienhardi.
Cormopsocus baleoi^[381]	Sp. nov	Valid	Hakim, Azar & Huang	Cretaceous	Burmese amber		A member of Trogiomorpha belonging to the family Cormopsocidae.
Cormopsocus neli^[382]	Sp. nov	Valid	Hakim, Azar & Huang in Hakim et al.	Cretaceous	Burmese amber		A member of Trogiomorpha belonging to the family Cormopsocidae.
Empherium^[383]	Gen. et sp. nov	Valid	Hakim, Huang & Azar	Late Cretaceous			A member of Trogiomorpha belonging to the family Empheriidae. Genus includes new species E. rasnitsyni.
Eolepinotus zherikhini	Sp. nov	Valid	Hakim, Huang & Azar	Late Cretaceous			A member of Trogiomorpha belonging to the family Trogiidae.
Heliadesdakruon^[384]	Gen. et sp. nov	Valid	Cumming & Le Tirant	Late Cretaceous (Cenomanian)	Burmese amber		A member of Trogiomorpha belonging to the family Archaeatropidae. The type species is H. morganae.
Longiglabellus^[385]	Gen. et 2 sp. nov	In press	Wang, Li & Yao in Wang et al.	Cretaceous	Burmese amber		A member of the family Cormopsocidae. Genus includes new species L. edentatus and L. pedhyalinus.
Palaeosiamoglaris hammanaensis^[386]	Sp. nov	In press	Hakim, Huang & Azar	Early Cretaceous (Barremian)	Lebanese amber		A member of Psocodea belonging to the family Prionoglarididae.
Palaeosiamoglaris hkamtiensis^[387]	Sp. nov	In press	Jouault et al.	Early Cretaceous	Hkamti amber		A member of Psocodea belonging to the family Prionoglarididae.
Psyllipsocus myanmarensis	Sp. nov	In press	Jouault et al.	Late Cretaceous (Cenomanian)	Burmese amber		A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.
Psyllipsocus yangi^[388]	Sp. nov	Valid	Liang & Liu	Cretaceous	Burmese amber		A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.
Stimulopsocus^[389]	Gen. et sp. nov	In press	Liang & Liu	Cretaceous	Burmese amber		A member of the family Cormopsocidae. Genus includes new species S. jiewenae.

Psocodean research

• Four bark lice carrying sand granules and organic material atop their back are described from the Cretaceous amber from Myanmar by Kiesmüller et al. (2021), who interpret this finding as the first fossil evidence of masking behaviour in Cretaceous representatives of Psocodea.^[390]

Clade Perlidea

Dermapterans

New Dermapteran

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aglyptoderma^[391]	Gen. et 2 sp. nov	Valid	Xiong et al.	Middle Jurassic	Jiulongshan Formation		A semenoviolid eodermapteran earwig. Genus includes the new species A. elongatum and A. cylindratum.
Aneuroderma^[392]	Gen. et sp. nov	Valid	Xiong, Engel & Ren in Xiong et al.	Middle Jurassic	Jiulongshan Formation		A member of the family Protodiplatyidae. The type species is A. oiodes.
Dacryoderma^[393]	Gen. et comb. nov	Valid	Engel	Early Jurassic (Sinemurian)	Charmouth Mudstone Formation		A dermapterid earwig. The type species is "Brevicula" teres Tihelka (2019).
Eminepygia^[394]	Gen. et sp. nov	Valid	Chen & Zhang	Cenomanian	Burmese amber		A pygidicranid earwig. Genus includes new species E. myanmarensis.
Sinopalaeodermata concavum	Sp. nov	Valid	Xiong, Engel & Ren in Xiong et al.	Middle Jurassic	Jiulongshan Formation		A member of the family Dermapteridae.

Plecoptera

New Plecopteran taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Balticonemoura^[395]	Gen. et sp. nov	In press	Chen	Eocene	Baltic amber	Europe (Baltic Sea region)	A nemourid stonefly. Genus includes new species B. bulbosus.
Billoperla^[396]	Gen. et sp. nov	In press	Chen & Xu	Cretaceous	Burmese amber		A perlid stonefly belonging to the family. Genus includes new species B. starki.
Isoperla baltica^[397]	Sp. nov	Valid	Jouault et al.	Eocene	Baltic amber	Europe (Baltic Sea regon)	A perlodid stonefly.

Other insects

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Anakitala^[398]	Gen. et sp. nov	Valid	Aristov	Late Permian			A mesorthopterid Eoblattidan Polyneopteran. The type species is A. intermittendia.
Baharellinus elegans	Sp. nov	Valid	Aristov	Late Permian			A member of Polyneoptera belonging to the group Eoblattida and the family Blattogryllidae.
Belmophenopterum rasnitsyni^[399]	Sp. nov	Valid	Storozhenko & Aristov	Triassic (Ladinian-Carnian)	Madygen Formation		A member of the family Mesorthopteridae.
Cabarzopterum^[400]	Gen. et sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation		A member of Grylloblattida of uncertain phylogenetic placement. Genus includes new species C. magnificus.
Delopterinus alsenzensis^[401]	Sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin		A miomopteran belonging to the family Palaeomanteidae.
Delopterum novokshonovi	Sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin		A miomopteran belonging to the family Palaeomanteidae.
Elmocladus^[402]	Gen. et comb. nov	Valid	Rasnitsyn & Aristov	Early Permian			A member of Polyneoptera belonging to the group Eoblattida and the family Tococladidae. The type species is "Opisthocladus" strictus Carpenter (1976).
Galloembia^[403]	Gen. et sp. nov	Valid	Falières, Engel & Nel	Earliest Eocene	Oise amber		A webspinner belonging to the family Embiidae. The type species is G. raholai.
Gnethoda odontophora^[404]	Sp. nov	In press	Lai et al.	Cretaceous	Burmese amber		A webspinner belonging to the family Clothodidae.
Kamopanorpa rasnitsyni^[405]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin		A member of the (stem-trichopteran or stem-amphiesmenopteran) family Microptysmatidae.
Kamopanorpa rotunda	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin		A member of the family Microptysmatidae.
Kamopanorpa sivchikovi	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin		A member of the family Microptysmatidae.
Liomopterites sperbersbachensis	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation		A member of Grylloblattida belonging to the family Liomopteridae.
Liomopterum fuscatum	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation		A member of Grylloblattida belonging to the family Liomopteridae.
Madygembia^[406]	Gen. et sp. nov	Valid	Aristov & Storozhenko	Triassic (Ladinian or Carnian)	Madygen Formation		A webspinner belonging to the family Alexarasniidae. The type species is M. rasnitsyni.
Magicivena^[407]	Fam. et gen. et 3 sp. nov	Valid	Yang et al.	Early Cretaceous	Yixian Formation		A member of Polyneoptera of uncertain phylogenetic placement, belonging to the new family Magicivenidae. The type species is M. sticta; genus also includes M. elegans and M. antennalis.
Multivena	Gen. et sp. nov	In press	Lai et al.	Cretaceous	Burmese amber		A webspinner belonging to the family Sorellembiidae. Genus includes new species M. curvivena.
Pictoborella	Gen. et sp. et comb. nov	Valid	Cui et al.	Early Permian			A member of Grylloblattida. Genus includes new species P. clara, as well as "Oborella" germanica Prokop et al. (2021).
Rasnalexia^[408]	Fam. et gen. et sp. nov	Valid	Gorochov	Triassic			A member of Embioptera belonging to the new family Rasnalexiidae. Genus includes new species R. rasnitsyni.
Uralioma thuringiensis	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation		A member of Grylloblattida belonging to the family Liomopteridae.

Other insect research

• A well-preserved full-body impression of an elongate winged insect strongly resembling stick insects is described from the Permian (Wordian) Pelitic Formation (France) by Logghe et al. (2021), who name a new ichnotaxon Phasmichnus radagasti.^[409]
• A dragonfly-type odonatopteran larva, preserving ancestral morphological characters maintained in mayflies but lost in modern odonatan larvae, is described from the Cretaceous Burmese amber by Haug, Müller & Haug (2021).^[410]

General research

• Revision of the fossil record of insects from the Triassic of China is published by Zhang et al. (2021).^[411]
• A study on the diversity of insect damage types in fossil plants from the Cretaceous (Albian to Cenomanian) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao et al. (2021).^[412]

Notes and References

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