2020 in paleontology explained

Plants

See main article: 2020 in paleobotany.

Sponges

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages
Endostoma stellata[1] Sp. novValidSenowbari-Daryan, Fürsich & RashidiJurassic (Callovian-Oxfordian)Qale-Dokhtar Limestone FormationA calcareous sponge belonging the family Endostomatidae.
Eoghanospongia[2] Gen. et sp. novValidBotting et al.Silurian (Telychian)A hexactinellid sponge. Genus includes new species E. carlinslowpensis. Announced in 2019; the final version of the article naming it was published in 2020.
Eudea maximaSp. novValidSenowbari-Daryan, Fürsich & RashidiJurassic (Callovian-Oxfordian)Qale-Dokhtar Limestone FormationA calcareous sponge belonging the family Endostomatidae.
IniquispongiaGen. et sp. novValidSenowbari-Daryan, Fürsich & RashidiJurassic (Callovian-Oxfordian)Qale-Dokhtar Limestone FormationA calcareous sponge belonging the family Endostomatidae. The type species is I. iranica.
Polyendostoma? irregularisSp. novValidSenowbari-Daryan, Fürsich & RashidiJurassic (Callovian-Oxfordian)Qale-Dokhtar Limestone FormationA calcareous sponge belonging the family Endostomatidae.
Polyendostoma? regularisSp. novValidSenowbari-Daryan, Fürsich & RashidiJurassic (Callovian-Oxfordian)Qale-Dokhtar Limestone FormationA calcareous sponge belonging the family Endostomatidae.
Preperonidella tabasensisSp. novValidSenowbari-Daryan, Fürsich & RashidiJurassic (Callovian-Oxfordian)Qale-Dokhtar Limestone FormationA calcareous sponge belonging the family Endostomatidae.
SeriespongiaGen. et sp. novValidSenowbari-Daryan, Fürsich & RashidiMiddle Jurassic (Callovian)Esfandiar Limestone FormationA calcareous sponge belonging the family Endostomatidae. The type species is S. iranica.
Shouzhispongia[3] Gen. et 2 sp. novIn pressBotting et al.Ordovician (Hirnantian)A rossellid sponge. Genus includes S. coronata and S. prodigia.
Spongia mantelli[4] Nom. novValidVan Soest, Hooper & ButlerCretaceousA replacement name for Spongia ramosa Mantell (1822).

Cnidarians

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Actinoseris riyadhensis[5] Sp. novValidGameil, El-Sorogy & Al-KahtanyLate Cretaceous (Campanian)ArumaA solitary coral. Announced in 2018; the final version of the article naming it was published in 2020.
Alichurastrea[6] Gen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. Genus includes new species A. major. Announced in 2020; the final version of the article naming it was published in 2021.
Alveopora kumadai[7] Sp. novValidNiko & SuzukiMiocene (Langhian)Katsuta GroupA species of Alveopora.
Amplexus gennarenensis[8] Sp. novValidLiao, Liang & LuoCarboniferous (Tournaisian)A rugose coral.
Anthracomedusa? hoferhauseri[9] Sp. novValidSzenteEarly TriassicWerfen FormationA box jellyfish.
Asteroseris arabicaSp. novValidGameil, El-Sorogy & Al-KahtanyLate Cretaceous (Campanian)ArumaA solitary coral. Announced in 2018; the final version of the article naming it was published in 2020.
Bowanophyllum ramosum[10] Sp. novValidWang, Percival & ZhenOrdovician (Katian)Malachis HillA rugose coral.
Carinthiaphyllum ramovsi[11] Sp. novValidKossovaya, Novak & WeyerPermian (Asselian)A rugose coral belonging to the family Geyerophyllidae.
Colligophyllum[12] Gen. et comb. novValidFedorowskiCarboniferous (Bashkirian)A rugose coral. The type species is "Lytvophyllum" dobroljubovae Vassilyuk (1960). Announced in 2020; the final version of the article naming it was published in 2021.
Cunnolites (Plesiocunnolites) riyadhensisSp. novValidGameil, El-Sorogy & Al-KahtanyLate Cretaceous (Campanian)ArumaA solitary coral. Announced in 2018; the final version of the article naming it was published in 2020.
Eohydnophora baingoinensis[13] Sp. novValidWang et al.Early CretaceousA stony coral.
EomicrophylliaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. Genus includes new species E. nalivkini. Announced in 2020; the final version of the article naming it was published in 2021.
Galliconularia[14] Gen. et comb. novValidVan Iten & LefebvreOrdovician (Tremadocian)Saint-ChinianA member of Conulariida. The type species is "Conularia" azaisi Thoral (1935).
GuembelastreomorphaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. Genus includes new species G. vinogradovi. Announced in 2020; the final version of the article naming it was published in 2021.
GurumdyniaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. Genus includes new species G. gracilis. Announced in 2020; the final version of the article naming it was published in 2021.
Hanagyroia[15] Gen. et sp. novValidWang et al.Early CambrianKuanchuanpuA medusozoan of uncertain phylogenetic placement, possibly representing an intermediate morphological type between scyphozoans and cubozoans. Genus includes new species H. orientalis.
Hemiagetiolites longiseptatusSp. novValidWang, Percival & ZhenOrdovician (Katian)Malachis HillA tabulate coral.
Heteroamphiastrea[16] Gen. et sp. novValidKołodziejEarly Cretaceous (Aptian)A stony coral belonging to the superfamily Heterocoenioidea and the family Carolastraeidae. Genus includes new species H. loeseri.
Heterostrotion huaqiaoense[17] Sp. novValidDenayer et al.Early CarboniferousA rugose coral
KrynkaphyllumGen. et 2 sp. novValidFedorowskiCarboniferous (Bashkirian)A rugose coral. The type species is K. multiplexum; genus also includes K. validum. Announced in 2020; the final version of the article naming it was published in 2021.
Martsaphyton[18] Gen. et sp. novValidTinn, Vinn & AinsaarOrdovician (Darriwilian)A member of Medusozoa of uncertain phylogenetic placement. The type species is M. moxi.
Michelinia flugeli[19] Sp. novValidNiko & BadpaCarboniferous (Bashkirian)Sardar FormationA tabulate coral belonging to the order Favositida and the family Micheliniidae.
Nancygyra[20] Gen. et sp. novIn pressBosellini & Stolarski in Bosellini et al.Eocene (Ypresian)A member of the family Euphylliidae. The type species is N. dissepimentata.
Neosyringaxon michelini[21] Sp. novValidWeyer & RohartDevonian (Frasnian)A rugose coral belonging to the family Petraiidae
Paramixogonaria wangyouensis[22] Sp. novValidLiao & LiangDevonian (Givetian)WenglaiA rugose coral.
PinacomorphaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. Genus includes new species P. apimelos. Announced in 2020; the final version of the article naming it was published in 2021.
Placophyllia baingoinensisSp. novValidWang et al.Early CretaceousA stony coral. Originally described as a species of Placophyllia, but subsequently transferred to the genus Sonoraphyllia.[23]
Placophyllia amnicaSp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A placophylliid coral. Announced in 2020; the final version of the article naming it was published in 2021.
Protokionophyllum feninoenseSp. novValidFedorowskiCarboniferous (Bashkirian)A rugose coral. Announced in 2020; the final version of the article naming it was published in 2021.
ProtostephanastreaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)An actinastraeid coral. Genus includes new species P. leveni. Announced in 2020; the final version of the article naming it was published in 2021.
PsenophylliaGen. et comb. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. The type species is "Cylindrosmilia" longa Melnikova (1989). Announced in 2020; the final version of the article naming it was published in 2021.
Rotiphyllum xinjiangenseSp. novValidLiao, Liang & LuoCarboniferous (Tournaisian)A rugose coral.
Sanidophyllum dubium[24] Sp. novValidYu et al.Devonian (Emsian)Mia LeA rugose coral belonging to the family Breviphyllidae. Announced in 2020; the final version of the article naming was published in 2021.
SedekastreaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A coral. Genus includes new species S. djalilovi. Announced in 2020; the final version of the article naming it was published in 2021.
Siphonophyllia khenifrense[25] Sp. novRodríguez, Said & Somerville in Rodríguez et al.Carboniferous (Viséan)Azrou-KhenifraA rugose coral belonging to the family Cyathopsidae
StylimorphaGen. et sp. novValidMelnikova & RoniewiczEarly Jurassic (probably Pliensbachian)A placophylliid coral. Genus includes new species S. kardjilgensis. Announced in 2020; the final version of the article naming it was published in 2021.
Stylina namcoensisSp. novValidWang et al.Early CretaceousA stony coral.
Stylostrotion houiSp. novValidDenayer et al.Carboniferous (Viséan)A rugose coral
Syringopora iranicaSp. novValidNiko & BadpaCarboniferous (Serpukhovian)Sardar FormationA tabulate coral belonging to the order Auloporida and the family Syringoporidae.

Research

Arthropods

See main article: 2020 in arthropod paleontology and 2020 in paleoentomology.

Bryozoans

Name Novelty Status Authors Age Type locality Country Notes Images
Anastomopora blankenheimensis[28] Sp. novValidErnstDevonian
Anastomopora minorSp. novValidErnstDevonian
Anomalotoechus parvus[29] Sp. novValidErnst, Bahrami & ParastDevonian (Famennian)BahramA member of Trepostomata belonging to the group Amplexoporina and to the family Atactotoechidae.
Asperopora sinensis[30] Sp. novValidErnst et al.Silurian (Telychian)HanchiatienA trepostome bryozoan.
Biforicula collinsi[31] Sp. novValidTaylorEarly Cretaceous (Albian)Gault
Cheethamia volgaensis[32] Sp. novValidKoromyslova & SeltserLate Cretaceous (Maastrichtian)
A member of Cheilostomata
Cribrilaria profunda[33] Sp. novValidRosso, Di Martino & OstrovskyPleistoceneA member of the family Cribrilinidae.
Dianulites altaicus[34] Sp. novValidKoromyslova & SennikovOrdovician (Sandbian)
A member of Esthonioporata.
Dyscritella kalmardensis[35] Sp. novValidErnst & GorgijCarboniferous (Pennsylvanian)Siliciclastic ImaghA member of Trepostomata belonging to the group Amplexoporina and to the family Dyscritellidae. Announced in 2019; the final version of the article naming it was published in 2020.
Dyscritella multiporataSp. novValidErnst & GorgijCarboniferous (Pennsylvanian)Siliciclastic ImaghA member of Trepostomata belonging to the group Amplexoporina and to the family Dyscritellidae. Announced in 2019; the final version of the article naming it was published in 2020.
Figularia spectabilisSp. novValidRosso, Di Martino & OstrovskyPleistoceneA member of the family Cribrilinidae.
Filites bakharevi[36] Sp. novValidMesentseva in Mesentseva & UdodovDevonian (Emsian)
Filites fragilisSp. novValidUdodov in Mesentseva & UdodovDevonian (Emsian)
Filites regularisSp. novValidMesentseva in Mesentseva & UdodovDevonian (Emsian)
Filites vulgarisSp. novValidUdodov in Mesentseva & UdodovDevonian (Emsian)
Glabrilaria transversocarinataSp. novValidRosso, Di Martino & OstrovskyPleistoceneA member of the family Cribrilinidae.
Hemiphragma insolitumSp. novValidKoromyslova & FedorovOrdovician (Dapingian)A trepostome bryozoan.
Microporella tanyae[37] Sp. novValidDi Martino, Taylor & GordonPlioceneYorktown
A member of the family Microporellidae.
Moorephylloporina parvulaSp. novValidErnst et al.Silurian (Telychian)HanchiatienA fenestrate bryozoan.
Parastenodiscus sinaiensis[38] Sp. novIn pressErnst et al.Carboniferous (Mississippian)A member of Trepostomata
Planopora[39] Gen. et sp. novValidKoromyslova & FedorovOrdovician (Dapingian)A bifoliate cystoporate. Genus includes new species P. volkhovensis.
Rhombopora aryaniSp. novValidErnst & GorgijCarboniferous (Pennsylvanian)Siliciclastic ImaghA member of Cryptostomata belonging to the group Rhabdomesina and to the family Rhomboporidae. Announced in 2019; the final version of the article naming it was published in 2020.
Taylorus patagonicus[40] Sp. novValidPérez et al.Early MioceneA member of the family Escharinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Trematopora jiebeiensisSp. novValidErnst et al.Silurian (Telychian)HanchiatienA trepostome bryozoan.
Trematopora tenuisSp. novValidErnst et al.Silurian (Telychian)HanchiatienA trepostome bryozoan.
ZefrehoporaGen. et sp. novValidErnst, Bahrami & ParastDevonian (Famennian)BahramA member of Trepostomata belonging to the group Amplexoporina and to the family Eridotrypellidae. The type species is Z. asynithis.

Brachiopods

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Altiplanotoechia[41] Gen. et sp. novValidColmenar in Colmenar & HodginOrdovicianUmachiriA polytoechioid brachiopod. Genus includes new species A. hodgini.
Beaussetithyris[42] Gen. et sp. novGaspard & CharbonnierLate Cretaceous (Santonian)A member of Rhynchonellida belonging to the family Cyclothyrididae. The type species is B. asymmetrica.
Biconvexiella saopauloensis[43] Sp. novIn pressSimões et al.Late PaleozoicTaciba
Bockeliena[44] Gen. et comb. novValidBaarliSilurian (Rhuddanian)A member of the family Atrypinidae; a new genus for "Atrypa" flexuosa Marr & Nicholson (1888).
Brevilamnulella minuta[45] Sp. novValidJin & BlodgettLate Ordovician
Chilcatreta lariojana[46] Sp. novValidLavié & BenedettoOrdovicianSuriA siphonotretid brachiopod. Announced in 2019; the final version of the article naming it was published in 2020.
Chinellirostra[47] Gen. et sp. novValidBaranov, Qiao & BlodgettDevonian (Givetian)A member of the family Stringocephalidae. Genus includes new species C. rara. Announced in 2020; the final version of the article naming was published in 2021.
ContortithyrisGen. et sp. novGaspard & CharbonnierLate Cretaceous (Santonian)MicrasterA member of Rhynchonellida belonging to the family Cyclothyrididae. The type species is C. thermae.
Cyclothyris cardiatelia[48] Sp. novIn pressBerrocal-Casero, Barroso-Barcenilla & JoralLate Cretaceous (Coniacian)A member of Rhynchonellida
Cyclothyris grimarginaSp. novGaspard & CharbonnierLate Cretaceous (Campanian)MicrasterA member of Rhynchonellida belonging to the family Cyclothyrididae
Cyclothyris nekvasilovae[49] Sp. novValidBerrocal-Casero, Joral & Barroso-BarcenillaLate Cretaceous (Cenomanian)A member of Rhynchonellida belonging to the family Cyclothyrididae. Announced in 2020; the final version of the article naming it was published in 2021.
Cyclothyris seguraiSp. novIn pressBerrocal-Casero, Barroso-Barcenilla & JoralLate Cretaceous (Coniacian)A member of Rhynchonellida
Dihelictera engerensisSp. novValidBaarliOrdovician/Silurian boundarySolvikA member of the family Atrypidae.
Dogdoa talyndzhensisSp. novValidBaranovEarly DevonianA member of Rhynchonellida.
Elliptoglossa kononovae[50] Sp. novValidSmirnova & ZhegalloDevonian (Famennian)A member of Lingulida.
EnriquetoechiaGen. et sp. novValidColmenar & HodginOrdovicianUmachiriA polytoechioid brachiopod. Genus includes new species E. umachiriensis.
Eoobolus incipiens[51] Sp. novIn pressZhang, Popov, Holmer & Zhang in Zhang et al.CambrianAjax Limestone
Dengying Formation
Mernmerna Formation
Wilkawillina Limestone
A member of Linguloidea.
Euroatrypa? sigridiSp. novValidBaarliOrdovician/Silurian boundarySolvikA member of the family Atrypinidae.
FamatinobolusGen. et sp. novValidLavié & BenedettoOrdovicianSuriAn obolid brachiopod. Genus includes new species F. cancellatum. Announced in 2019; the final version of the article naming it was published in 2020.
Germanoplatidia[52] Gen. et comb. novValidDulai & Von der HochtOligocene (Chattian)A member of Terebratulida belonging to the family Platidiidae; a new genus for "Terebratula" pusilla Philippi (1843).
Gotatrypa vettrensisSp. novValidBaarliOrdovician/Silurian boundarySolvikA member of the family Atrypidae.
Hassanispirifer[53] Gen. et sp. novValidGarcia-Alcalde & El HassaniDevonian (Givetian)TaboumakhloufA member of Spiriferida belonging to the family Xenomartiniidae. The type species is H. africanus.
Holynetes? mzerrebiensisSp. novValidGarcia-Alcalde & El HassaniDevonian (Givetian)AhrerouchA member of Chonetidina belonging to the family Anopliidae.
Imbriea[54] Nom. novValidReilyDevonianA member of Orthotetida belonging to the family Areostrophiidae; a replacement name for Orthopleura Imbrie (1959).
Kafirnigania jorali[55] Sp. novIn pressBerrocal-CaseroLate Cretaceous (Coniacian)A member of Terebratulida.
Kafirnigania massiliensisSp. novIn pressBerrocal-CaseroLate Cretaceous (Coniacian)
A member of Terebratulida.
Kirkidium canberrense[56] Sp. novValidStruszSilurian (Wenlock)CanberraA member of Pentamerida belonging to the family Pentameridae.
Lambdarina winklerprinsi[57] Sp. novValidVoldman et al.Carboniferous (Pennsylvanian)San Emiliano
Levitusia elongata[58] Sp. novValidTazawaCarboniferous (Viséan)A member of Productidina belonging to the family Leioproductidae.
Lingulellotreta yuanshanensis[59] Sp. novValidZhang et al.Cambrian
Linnaeocaninella[60] Nom. novValidHernández Middle PermianLengwuA replacement name for Caninella Liang (1990)
Linnarssonia sapushanensis[61] Sp. novValidDuan et al.Cambrian Stage 4WulongqingAn acrotretoid brachiopod.
Lithobolus limbatumSp. novValidLavié & BenedettoOrdovicianSuriAn obolid brachiopod. Announced in 2019; the final version of the article naming it was published in 2020.
Mishninia[62] Gen. et sp. novValidBaranovEarly DevonianThe type species is M. nodosa
Neobolus wulongqingensis[63] Sp. novValidZhang, Strotz, Topper & Brock in Zhang et al.Cambrian Stage 4WulongqingA member of Lingulida belonging to the family Neobolidae. Many specimens had tubeworm-like kleptoparasites attached to their shells.
Neochonetes (Sommeriella) longa[64] Sp. novValidWu et al.Permian (Changhsingian)Luokeng
Neochonetes (Sommeriella) transversa Sp. novValidWu et al.Permian (Changhsingian)Luokeng
Nucleatina anotiaSp. novIn pressBerrocal-CaseroLate Cretaceous (Coniacian)
?		A member of Terebratulida.
Nucleatina arcanaSp. novIn pressBerrocal-CaseroLate Cretaceous (Coniacian)A member of Terebratulida.
Nucleatina barrosoiSp. novIn pressBerrocal-CaseroLate Cretaceous (Coniacian)A member of Terebratulida.
Orbiculoidea katzeri[65] Sp. novIn pressCorrêa & RamosDevonian (Lochkovian)Manacapuru
Orbiculoidea xinguensisSp. novIn pressCorrêa & RamosDevonian (Lochkovian)Manacapuru
Palaeotreta[66] Gen. et sp. et comb. novValidZhang et al.Cambrian Series 2ShuijingtuoA member of the family Acrotretidae. The type species is P. shannanensis; genus also includes "Eohadrotreta" zhujiahensis Li & Holmer (2004).
Paragilledia[67] Gen. et sp. novValidShi, Waterhouse & LeeEarly PermianPebbley BeachA member of Terebratulida belonging to the family Gillediidae. Genus includes new species P. kioloaensis.
Paramickwitzia[68] Gen. et sp. novValidPan et al.Cambrian Series 2XinjiA stem-brachiopod belonging to the group Mickwitziidae. Genus includes new species P. boreussinaensis. Announced in 2019; the final version of the article naming it was published in 2020.
Plectatrypa rindiSp. novValidBaarliOrdovician/Silurian boundarySolvikA member of the family Atrypinidae.
Plicarmus[69] Gen. et sp. novValidClaybourn et al.Cambrian Stage 4Byrd GroupAntarcticaA member of Lingulata. Genus includes new species P. wildi.
Pomatotrema laubacheriSp. novValidColmenar & HodginOrdovicianUmachiri
RhinatrypaGen. et comb. novValidBaarliOrdovician/Silurian boundarySolvikA member of the family Atrypidae. The type species is "Plectatrypa" henningsmoeni Boucot & Johnson (1967).
Rhipidium oepikiSp. novValidStruszSilurian (Wenlock)CanberraA member of Pentamerida belonging to the family Pentameridae.
Spinocarinifera qilinzhaiensis[70] Sp. novValidNie et al.Carboniferous (Tournaisian)Tangbagou Formation
Stringocephalus sinensisSp. novValidBaranov, Qiao & BlodgettDevonian (Givetian)A member of the family Stringocephalidae. Announced in 2020; the final version of the article naming was published in 2021.
Tabellina laseroniSp. novValidShi, Waterhouse & LeeEarly PermianPebbley BeachAn ingelarelloidean brachiopod belonging to the family Notospiriferidae.
Tapuritreta gribovensis[71] Sp. novValidHolmer et al.Cambrian (Guzhangian)Karpinsk Formation
A member of the family Acrotretidae.
Tcherskidium tenuicostatusSp. novValidJin & BlodgettLate Ordovician
Thomasaria bultynckiSp. novValidGarcia-Alcalde & El HassaniDevonian (Givetian)AhrerouchA member of Spiriferida belonging to the family Thomasariidae.
Vagrania naanchanensisSp. novValidBaranovEarly DevonianA member of Atrypida.
Verchojania abramovi[72] Sp. novValidMakoshinLate CarboniferousA member of Productida
Wahwahlingula? pankovensisSp. novValidHolmer et al.Cambrian (Guzhangian)Karpinsk Formation
A member of Linguloidea belonging to the family Zhanatellidae.
Woodwardirhynchia pontemdiaboliSp. novIn pressBerrocal Casero, Barroso Barcenilla & JoralLate Cretaceous (Coniacian)A member of Rhynchonellida
YangirostraGen. et sp. novValidBaranov, Qiao & BlodgettDevonian (Givetian)A member of the family Stringocephalidae. Genus includes new species Y. asiatica. Announced in 2020; the final version of the article naming was published in 2021.

Research

Molluscs

See main article: 2020 in paleomalacology.

Echinoderms

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Abertella carlsoni[76] Sp. novValidOsborn, Portell & MooiMiocene United States
A sea urchin.
Abludoglyptocrinus steinheimerae[77] Sp. novValidCole et al.Ordovician (Katian)Brechin Lagerstätte
Bobcaygeon & Verulam
A monobathrid crinoid.
Aenigmaticumcrinus[78] Gen. et sp. novValidSchefflerDevonianBelénA crinoid belonging to the group Dimerocrinitacea. Genus includes new species A. rochacamposi.
Aerliceaster[79] Gen. et sp. novValidBlake, Gahn & GuensburgOrdovician (Floian)Garden City
A starfish. Genus includes new species A. nexosus.
Alkaidia megaungula[80] Sp. novValidEwin & GaleEarly Cretaceous (Barremian)TabaA starfish belonging to the family Terminasteridae.
Arceoaster[81] Gen. et sp. novValidBlake & SprinkleSilurianHunton Group United States
A starfish belonging to the family Hudsonasteridae. Genus includes new species A. hintei.
Aszulcicrinus[82] Gen. et sp. novValidHagdornMiddle Triassic (Anisian)GogolinA crinoid belonging to the group Articulata and the family Dadocrinidae. The type species is A. pentebrachiatus.
Brissopsis hoffmaniSp. novValidOsborn, Portell & MooiMiocene United States
A sea urchin.
Bronthaster[83] Gen. et sp. novIn pressJell & CookCarboniferous (Namurian)Yagon SiltstoneA brittle star belonging to the family Protasteridae. Genus includes new species B. retus.
Calclyra bifida[84] Sp. novValidPabst & HerbigCarboniferous (Serpukhovian)Genicera SpainA brittle star belonging to the group Oegophiurida and the family Calclyridae.
Clypeaster petersonorumSp. novValidOsborn, Portell & MooiMiocene United States
A species of Clypeaster.
Comptonia bretoni[85] Sp. novValidGaleEarly Cretaceous (Aptian)AtherfieldA starfish
Coulonia caseyiSp. novValidGaleEarly Cretaceous (Aptian)AtherfieldAn astropectinid starfish
Cyclogrupera[86] Gen. et sp. novTorres-Martínez, Villanueva-Olea & Sour-TovarPermian (Asselian‒Sakmarian)GruperaA crinoid belonging to the family Cyclomischidae. The type species is C. minor.
Discocrinus africanus[87] Sp. novValidGaleLate Cretaceous (Cenomanian)Aït LamineA crinoid belonging to the group Articulata and the family Roveacrinidae.
Discometra luberonensis[88] Sp. novValidEléaume, Roux & PhilippeMiocene (Burdigalian)A feather star belonging to the family Himerometridae.
Drepanocrinus wardorumSp. novValidGaleLate Cretaceous (Cenomanian)
A crinoid belonging to the group Articulata and the family Roveacrinidae
Durhamicystis[89] Gen. et sp. novValidZamora, Sprinkle & SumrallOrdovician (Sandbian)Chambersburg
A member of Eocrinoidea belonging to the family Rhipidocystidae. The type species is D. americana.
Encrinaster alsbachensis[90] Sp. novValidMüller & HahnEarly DevonianA brittle star.
Enodicalix[91] Gen. et comb. novValidPaul & Gutiérrez-MarcoOrdovician SpainA member of Diploporita belonging to the family Aristocystitidae. The type species is "Calix" inornatus Meléndez (1958).
Eoastropecten[92] Gen. et sp. novValidGaleLate Triassic (Carnian) ChinaA starfish belonging to the family Astropectinidae. Genus includes new species E. sechuanensis.
Euglyphocrinus cristagalliSp. novValidGaleEarly Cretaceous (Albian)

A crinoid belonging to the group Articulata and the family Roveacrinidae
Euglyphocrinus jacobsaeSp. novValidGaleLate Cretaceous (Cenomanian)
A crinoid belonging to the group Articulata and the family Roveacrinidae
Euglyphocrinus truncatusSp. novValidGaleLate Cretaceous (Cenomanian)
A crinoid belonging to the group Articulata and the family Roveacrinidae
Euglyphocrinus worthensisSp. novValidGaleEarly Cretaceous (Albian)

A crinoid belonging to the group Articulata and the family Roveacrinidae
Euptychocrinus longipinnulus[93] Sp. novValidFearnhead et al.Silurian (Telychian)Pysgotwr GritsA camerate crinoid
Eutaxocrinus ariunai[94] Sp. novValidWaters et al.Devonian (Famennian)Samnuuruul Formation MongoliaA crinoid. Announced in 2020; the final version of the article naming was published in 2021.
Eutaxocrinus sersmaaiSp. novValidWaters et al.Devonian (Famennian)Samnuuruul Formation MongoliaA crinoid. Announced in 2020; the final version of the article naming was published in 2021.
FenestracrinusGen. et sp. novValidGaleLate Cretaceous (Cenomanian)Aït LamineA crinoid belonging to the group Articulata and the family Roveacrinidae. The type species is F. oculifer.
Fernandezaster whisleriSp. novValidOsborn, Portell & MooiPliocene United States
A sea urchin.
FloricyclocionGen. et sp. novTorres-Martínez, Villanueva-Olea & Sour-TovarPermian (Asselian‒Sakmarian)GruperaA crinoid belonging to the family Floricyclidae. The type species is F. heteromorpha.
Gagaria hunteraeSp. novValidOsborn, Portell & MooiMiocene United States
A sea urchin.
Genocidaris oyeniSp. novValidOsborn, Portell & MooiPliocene United States
A sea urchin.
Heterobrissus lubellii[95] Sp. novValidBorghi & StaraLate Oligocene-early MioceneA heart urchin.
Holocrinus qingyanensis[96] Sp. novValidStillerMiddle Triassic (Anisian)A crinoid belonging to the family Holocrinidae. Announced in 2019; the final version of the article naming it was published in 2020.
Isocrinus (Chladocrinus) covuncoensis[97] Sp. novValidLazo et al.Early Cretaceous (Valanginian)AgrioA crinoid.
Isocrinus (Chladocrinus) pehuenchensisSp. novValidLazo et al.Early Cretaceous (Hauterivian)AgrioA crinoid.
KolatasterGen. et sp. novValidBlake, Gahn & GuensburgOrdovician (Sandian)Mifflin
A starfish. Genus includes new species K. perplexus.
Lebenharticrinus quinvigintensisSp. novValidGaleLate Cretaceous (Cenomanian)Aït LamineA crinoid belonging to the group Articulata and the family Roveacrinidae
Lebenharticrinus zittiSp. novValidGaleLate Cretaceous (Cenomanian)Aït LamineA crinoid belonging to the group Articulata and the family Roveacrinidae
Linguaserra heidiiSp. novValidPabst & HerbigCarboniferous (Tournaisian to Serpukhovian)Genicera
Heiligenhaus		 Germany
Spain		A member of Ophiocistioidea belonging to the family Linguaserridae.
Lovenia kerneriSp. novValidOsborn, Portell & MooiPliocene United States
A species of Lovenia.
Maestratina[98] Gen. et comb. novValidForner i Valls & Saura VilarEarly Cretaceous (Aptian)Forcall FormationA sea urchin belonging to the group Arbacioida and the family Arbaciidae. The type species is "Cotteaudia" royoi Lambert (1928).
Magnasterella[99] Gen. et comb. novIn pressFraga & VegaDevonian (Frasnian)Ponta GrossaA starfish belonging to the group Euaxosida; a new genus for "Echinasterella" darwini Clarke (1913).
Marginix notatusSp. novIn pressFraga & VegaDevonian (Frasnian)Ponta GrossaA brittle star
MeperocrinusGen. et sp. novValidSchefflerDevonianIclaA crinoid belonging to the family Emperocrinidae. Genus includes new species M. angelina.
MongoliacrinusGen. et sp. novValidWaters et al.Devonian (Famennian)Samnuuruul Formation MongoliaA crinoid belonging to the family Acrocrinidae. Genus includes new species M. minjini. Announced in 2020; the final version of the article naming was published in 2021.
Odontaster tabaensisSp. novValidEwin & GaleEarly Cretaceous (Barremian)TabaA starfish, a species of Odontaster.
Ophiacantha oceani[100] Sp. novValidNumberger-Thuy & ThuyPliocene to Pleistocene (Piacenzian to Gelasian)A brittle star belonging to the family Ophiacanthidae.
Ophiomitrella floorae[101] Sp. novValidThuy, Numberger-Thuy & GaleLate Cretaceous (Maastrichtian)MaastrichtAn ophiacanthid brittle star.
Paragonaster felli[102] Sp. novValidStevensEarly CretaceousA starfish.
ParanasterGen. et comb. novIn pressFraga & VegaDevonian (Emsian)Ponta GrossaA starfish belonging to the group Euaxosida. Genus includes new species P. crucis.
Pararchaeocrinus kiddiSp. novValidCole et al.Ordovician (Katian)Brechin Lagerstätte
Bobcaygeon & Verulam
A diplobathrid crinoid.
Peckicrinus[103] Gen. et comb. novValidGale in Gale et al.Early Cretaceous (Albian)Duck Creek
(
)		A crinoid belonging to the family Roveacrinidae. The type species is "Poecilocrinus" porcatus Peck (1943). Announced in 2020; the final version of the article naming it was published in 2021.
Pegoasterella[104] Gen. et sp. novValidBlake & KonieckiLate OrdovicianBromide
Guttenberg
(
)		A starfish belonging to the family Urasterellidae. Genus includes new species P. pompom.
Periglyptocrinus astricusSp. novValidCole et al.Ordovician (Katian)Brechin Lagerstätte
Bobcaygeon & Verulam
A monobathrid crinoid.
Periglyptocrinus kevinbrettiSp. novValidCole et al.Ordovician (Katian)Brechin Lagerstätte
Bobcaygeon & Verulam
A monobathrid crinoid.
Periglyptocrinus mcdonaldiSp. novValidCole et al.Ordovician (Katian)Brechin Lagerstätte
Bobcaygeon & Verulam
A monobathrid crinoid.
Periglyptocrinus silvosusSp. novValidCole et al.Ordovician (Katian)Brechin Lagerstätte
Bobcaygeon & Verulam
A monobathrid crinoid.
Plotocrinus molineuxaeSp. novValidGale in Gale et al.Early Cretaceous (Albian)Goodland
A crinoid belonging to the family Roveacrinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Plotocrinus rashallaeSp. novValidGale in Gale et al.Early Cretaceous (Albian)Goodland

A crinoid belonging to the family Roveacrinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Plotocrinus reidiSp. novValidGale in Gale et al.Early Cretaceous (Albian)Kiamichi
A crinoid belonging to the family Roveacrinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Psammaster[105] Gen. et comb. novValidFau et al.Late Jurassic (Tithonian)Grès des OiesA starfish belonging to the group Forcipulatida. The type species is "Ophidiaster" davidsoni de Loriol & Pellat (1874).
Rhyncholampas meansiSp. novValidOsborn, Portell & MooiPleistocene United States
A sea urchin.
Roveacrinus gladiusSp. novValidGaleLate Cretaceous (Cenomanian)
A crinoid belonging to the group Articulata and the family Roveacrinidae
Roveacrinus morganaeSp. novValidGale in Gale et al.Early Cretaceous (Albian)Pawpaw
A crinoid belonging to the family Roveacrinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Roveacrinus proteusSp. novValidGale in Gale et al.Early Cretaceous (Albian)Pawpaw
A crinoid belonging to the family Roveacrinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Roveacrinus solisoccasumSp. novValidGaleEarly Cretaceous (Albian)

A crinoid belonging to the group Articulata and the family Roveacrinidae
Schoenaster carterensis[106] Sp. novValidHarris, Ettensohn & Carnahan-JarvisCarboniferous (Chesterian)Slade
A brittle star
Seifenia[107] Gen. et sp. novValidMüller & HahnEarly DevonianSeifenA member of Edrioasteroidea. The type species is S. ostara.
Spiracarneyella[108] Gen. et sp. novValidSumrall & PhelpsOrdovician (Katian)Point Pleasant
(
)		A carneyellid edrioasteroid. Genus includes new species S. florencei.
Streptoiocrinus[109] Gen. novValidRozhnovOrdovician

A crinoid belonging to the group Disparida.
Styracocrinus rimaferaSp. novValidGaleLate Cretaceous (Cenomanian)
A crinoid belonging to the group Articulata and the family Roveacrinidae
Styracocrinus thomasaeSp. novValidGale in Gale et al.Early Cretaceous (Albian)Goodland
A crinoid belonging to the family Roveacrinidae. Announced in 2020; the final version of the article naming it was published in 2021.
Tallinnicrinus[110] Gen. et sp. novValidCole, Ausich & WilsonOrdovician (Hirnantian)An anthracocrinid diplobathrid crinoid. Genus includes new species T. toomae.
Tollmannicrinus leidapoensisSp. novValidStillerMiddle Triassic (Anisian)A crinoid. Announced in 2019; the final version of the article naming it was published in 2020.
TuberocrinusGen. et sp. novValidSchefflerDevonianBelénA crinoid belonging to the group Dimerocrinitacea. Genus includes new species T. lapazensis.
Vaquerosella perrillatae[111] Sp. novValidMartínez Melo & Alvarado OrtegaMioceneSan IgnacioA sand dollar belonging to the family Echinarachniidae

Research

Conodonts

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Ancyrognathus minjini[120] Sp. novValidSuttner et al.Late DevonianBaruunhuuraiAnnounced in 2019; the final version of the article naming it was published in 2020.
Baltoniodus norrlandicus denticulatus[121] Subsp. novValidDzikOrdovician (Darriwilian)Announced in 2019; the final version of the article naming it was published in 2020.
Belodina watsoni[122] Sp. novValidZhenOrdovician (Darriwilian)
Bipennatus hemilevigatus[123] Sp. novValidLu & KönigshofDevonian (Eifelian)BeiliuAnnounced in 2019; the final version of the article naming it was published in 2020.
Bipennatus planusSp. novValidLu & KönigshofDevonian (Eifelian)BeiliuAnnounced in 2019; the final version of the article naming it was published in 2020.
Diplognathodus benderi[124] Sp. novValidHu et al.Carboniferous (Bashkirian–Moscovian boundary)
Erraticodon neopatu[125] Sp. novValidZhen in Zhen et al.OrdovicianWillaraAnnounced in 2020; the final version of the article naming it was published in 2021.
Gladigondolella laii[126] Sp. novIn pressChen in Chen et al.Early Triassic
Idiognathodus fengtingensis[127] Sp. novValidQi et al.Carboniferous (Kasimovian–Gzhelian boundary)
Idiognathodus luodianensisSp. novValidQi et al.Carboniferous (Kasimovian–Gzhelian boundary)
Idiognathodus naqingensisSp. novValidQi et al.Carboniferous (Kasimovian–Gzhelian boundary)
Idiognathodus naraoensisSp. novValidQi et al.Carboniferous (Kasimovian–Gzhelian boundary)
Latericriodus guangnanensis[128] Sp. novIn pressLu & Valenzuela-Ríos in Lu et al.Devonian (Emsian)DaliantangA member of Prioniodontida belonging to the family Icriodontidae.
Misikella kolarae[129] Sp. novValidKarádi et al.Late TriassicAnnounced in 2019; the final version of the article naming it was published in 2020.
Pachycladina rendona[130] Sp. novIn pressWu & Ji in Wu et al.Early TriassicAn ellisonid conodont.
Palmatolepis subperlobata tatarica[131] Nom. novValidOvnatanova & GatovskyDevonian (Famennian)Prikazanskaya Formation
A replacement name for Palmatolepis subperlobata helmsi Ovnatanova (1976). The subspecies was subsequently raised to the rank of a separate species by Ovnatanova & Kononova (2023).[132]
Paullella omanensisSp. novIn pressChen in Chen et al.Early Triassic
Polygnathus nalaiensisSp. novValidLu & KönigshofDevonian (Eifelian)BeiliuAnnounced in 2019; the final version of the article naming it was published in 2020.
Rossodus? boothiaensis[133] Sp. novValidZhangTurner Cliffs
Scalpellodus percivaliSp. novValidZhenOrdovician (Darriwilian)
Scythogondolella dolosa[134] Sp. novValidBondarenko & PopovEarly Triassic
Siphonodella leiosa[135] Sp. novIn pressSouquet, Corradini & GirardCarboniferous (Tournaisian)
Streptognathodus nemyrovskaeSp. novValidQi et al.Carboniferous (Gzhelian)
Streptognathodus zhihaoiSp. novValidQi et al.Carboniferous (Gzhelian)
Tortodus dodoensis[136] Sp. novValidGouwy, Uyeno & McCrackenDevonian (Givetian)Announced in 2019; the final version of the article naming it was published in 2020.
Trapezognathus pectinatusSp. novValidDzikOrdovician (Darriwilian)Announced in 2019; the final version of the article naming it was published in 2020.
Zieglerodina petrea[137] Sp. novValidHušková & SlavíkSilurian/Devonian boundaryPrague SynformAnnounced in 2019; the final version of the article naming it was published in 2020.

Research

Fishes

See main article: 2020 in paleoichthyology.

Amphibians

See main article: 2020 in amphibian paleontology.

Reptiles

See main article: 2020 in reptile paleontology.

Synapsids

Non-mammalian synapsids

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Agudotherium[140] Gen. et sp. novValidStefanello et al.Late TriassicCandeláriaA non-mammaliaform prozostrodontian cynodont. Genus includes new species A. gassenae.
BohemiclavulusGen. et comb. novValidSpindler, Voigt & FischerCarboniferous (Gzhelian)SlanýA member of the family Edaphosauridae; a new genus for "Naosaurus" mirabilis Fritsch (1895). Announced in 2019; the final version of the article naming it was published in 2020.
Caodeyao[141] Gen. et sp. novValidLiu & AbdalaLate PermianNaobaogouA therocephalian. Genus includes new species C. liuyufengi.
Chiniquodon omaruruensis[142] Sp. novValidMocke, Gaetano & AbdalaTriassicOmingonde
Dendromaia[143] Gen. et sp. novValidMaddin, Mann & HebertCarboniferous
A member of Varanopidae. Genus includes new species D. unamakiensis. Announced in 2019; the final version of the article naming it was published in 2020.
Etjoia[144] Gen. et sp. novValidHendrickx et al.Triassic (Ladinian/Carnian)OmingondeA traversodontid cynodont. Genus includes new species E. dentitransitus.
Hypselohaptodus[145] Gen. et comb. novValidSpindlerPermian (Cisuralian)KenilworthAn early member of Sphenacodontia; a new genus for "Haptodus" grandis. Announced in 2019; the final version of the article naming it was published in 2020.
Inditherium[146] Gen. et sp. novValidBhat, Ray & DattaLate TriassicTikiA dromatheriid cynodont. Genus includes new species I. floris.
Kalaallitkigun[147] Gen. et sp. novValidSulej et al.Late Triassic (Norian)Fleming FjordAn early member of Mammaliaformes, possibly a member of Haramiyida. Genus includes new species K. jenkinsi.
Kataigidodon[148] Gen. et sp. novValidKligman et al.Late TriassicChinle
A non-mammalian eucynodont. Genus includes new species K. venetus.
Kenomagnathus[149] Gen. et sp. novValidSpindlerCarboniferous (late Pennsylvanian)Rock Lake Shale Mb, Stanton
An early member of Sphenacodontia. The type species is K. scottae.
Martensius[150] Gen. et sp. novValidBerman et al.Permian (Artinskian)TambachA member of Caseidae. The type species is M. bromackerensis.
Nshimbodon[151] Gen. et sp. novValidHuttenlocker & SidorLate PermianMadumabisa MudstoneA basal cynodont, probably a member of the family Charassognathidae. Genus includes new species N. muchingaensis.
Polonodon[152] Gen. et sp. novValidSulej et al.Late Triassic (Carnian)A non-mammaliaform eucynodont. Genus includes new species P. woznikiensis. Announced in 2018; the final version of the article naming it was published in 2020.
Remigiomontanus[153] Gen. et sp. novValidSpindler, Voigt & FischerCarboniferous–Permian transitionSaar–NaheA member of the family Edaphosauridae. Genus includes new species R. robustus. Announced in 2019; the final version of the article naming it was published in 2020.
Rewaconodon indicusSp. novValidBhat, Ray & DattaLate TriassicTikiA dromatheriid cynodont.
Taoheodon[154] Gen. et sp. novValidLiuLate PermianSunjiagou FormationA dicynodontoid dicynodont. Genus includes new species T. baizhijuni.
Theroteinus jenkinsi[155] Sp. novValidWhiteside & DuffinLate Triassic (Rhaetian)A haramiyidan mammaliaform. Announced in 2020; the final version of the article naming it was published in 2021.
TikiodonGen. et sp. novValidBhat, Ray & DattaLate TriassicTikiA mammaliamorph cynodont. Genus includes new species T. cromptoni.

Research

Mammals

Other animals

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Aladraco kirchhainensis[176] Sp. novValidGeyer & MalinkyCambrian (Miaolingian)Delitzsch–Torgau–DoberlugA member of Hyolitha. Announced in 2019; the final version of the article naming it was published in 2020.
Armilimax[177] Gen. et sp. novValidKimmig & SeldenCambrian (Wuliuan)Spence Shale
A shell-bearing animal of uncertain phylogenetic placement. Genus includes new species A. pauljamisoni. Announced in 2020; the final version of the article naming it was published in 2021.
Avitograptus akidomorphus[178] Sp. novValidMuir et al.Ordovician (Hirnantian)WenchangA graptolite.
BizeticyathusGen. et comb. novValidKruse & DebrenneCambrianA member of Archaeocyatha. Genus includes B. carmen (Carmen & Carmen, 1937).
Canadiella[179] Gen. et comb. novValidSkovsted et al.CambrianMural
Rosella		A tommotiid belonging to the family Kennardiidae. The type species is "Lapworthella" filigrana Conway Morris & Fritz (1984).
Collinsovermis[180] Gen. et sp. novValidCaron & AriaCambrian (Wuliuan)Burgess Shale
A luolishaniid lobopodian. Genus includes new species C. monstruosus.
Cordaticaris[181] Gen. et sp. novIn pressSun, Zeng & ZhaoCambrian (Drumian)ZhangxiaA member of Radiodonta belonging to the family Hurdiidae. Genus includes new species C. striatus.
Cornulites baranovi[182] Sp. novValidVinn & ToomSilurian (Přidoli)OhesaareA member of Cornulitida.
Dahescolex[183] Gen. et sp. novValidShao et al.Cambrian (Fortunian)KuanchuanpuAn animal which might be a stem-lineage derivative of Scalidophora. Genus includes new species D. kuanchuanpuensis. Announced in 2019; the final version of the article naming it was published in 2020.
Dakorhachis[184] Gen. et sp. novValidConway Morris et al.Cambrian (Guzhangian)Weeks
An animal of uncertain phylogenetic placement, possibly a stem-group member of the Gnathifera. Genus includes new species D. thambus.
Dannychaeta[185] Gen. et sp. novValidChen et al.Early CambrianCanglangpuA crown annelid, probably a relative of the families Magelonidae and Oweniidae. Genus includes new species D. tucolus.
Degeletticyathus dailyiSp. novValidKruse & DebrenneCambrianA member of Archaeocyatha.
"Dictyonema" khadijae[186] Sp. novIn pressGutiérrez Marco, Muir & MitchellLate OrdovicianA graptolite
"Dictyonema" villasiSp. novIn pressGutiérrez Marco, Muir & MitchellLate OrdovicianA graptolite
Gyaltsenglossus[187] Gen. et sp. novValidNanglu, Caron & CameronCambrianStephen
A member of the stem group of Hemichordata. The type species is G. senis.
Herpetogaster haiyanensis[188] Sp. novYang et al.Cambrian Stage 3Chiungchussu
HillaecyathusGen. et comb. novValidKruse & DebrenneCambrianA member of Archaeocyatha. Genus includes H. contractus (Hill, 1965).
Ikaria[189] Gen. et sp. novValidEvans et al.EdiacaranAn early bilaterian. Genus includes new species I. wariootia.
Korenograptus selectus[190] Sp. novIn pressChen in Chen et al.Late OrdovicianA graptolite
Kylinxia[191] Gen. et sp. novValidZeng, Zhao & Huang in Zeng et al.Early CambrianA transitional euarthropod that bridges radiodonts and true arthropods. Genus includes new species K. zhangi.
Lenzograptus[192] Nom. novIn pressLoydellSilurian (Ludlow)
A graptolite; a replacement name for Lenzia Rickards & Wright (1999).
Longxiantheca[193] Gen. et sp. novValidLi in Li et al.Cambrian Stages 3–4XinjiA member of Hyolitha belonging to the group Orthothecida. The type species is L. mira.
Maxdebrennius[194] Gen. et sp. novValidKruse & DebrenneCambrianA member of Archaeocyatha. Genus includes new species M. mimus.
Microconchus cravenensis[195] Sp. novValidZatoń & MundyCarboniferous (Mississippian)Cracoe Limestone
Malham		A member of Microconchida.
Microconchus maya[196] Sp. novValidHeredia-Jiménez et al.Permian (Roadian)Paso HondoA member of Microconchida.
Monograptus hamulus[197] Sp. novValidSaparin et al.Silurian (Llandovery)Co ToA graptolite
Neodiplograptus mandalayensisSp. novIn pressChen in Chen et al.Late OrdovicianA graptolite
Nochoroicyathus ordinariusSp. novValidKruse & DebrenneCambrianA member of Archaeocyatha.
Nochoroicyathus sublimusSp. novValidKruse & DebrenneCambrianA member of Archaeocyatha.
Paranacyathus arboreusSp. novValidKruse & DebrenneCambrianA member of Archaeocyatha.
Pontagrossia[198] Gen. et sp. novValidChahud & FairchildDevonian (Emsian)Ponta GrossaAn invertebrate of uncertain phylogenetic placement. The type species is P. reticulata.
Porocoscinus eurysSp. novValidKruse & DebrenneCambrianA member of Archaeocyatha.
Pristiograptus paradoxus[199] Sp. novIn pressLoydell & WalasekSilurian (Telychian)A graptolite
StictocyathusGen. et sp. novValidKruse & DebrenneCambrianA member of Archaeocyatha. Genus includes new species S. cavus.
Subtumulocyathellus satusSp. novValidKruse & DebrenneCambrianA member of Archaeocyatha.
Torquigraptus loveridgeiSp. novIn pressLoydell & WalasekSilurian (Telychian)A graptolite
Torquigraptus wilsoni[200] Sp. novValidLoydellSilurian (Telychian)A graptolite
Toscanisoma[201] Gen. et 2 sp. novValidWendtLate Triassic (Carnian)San CassianoA member of Ascidiacea. The type species is T. multipartitum; genus also includes T. triplicatum.
Utahscolex[202] Gen. et comb. novValidWhitaker et al.Cambrian (Wuliuan)Spence
A palaeoscolecid; a new genus for "Palaeoscolex" ratcliffei Robison (1969)
Vermilituus[203] Gen. et sp. novValidLi et al.Cambrian Stage 3ChiungchussuA small, encrusting tubular protostomian, preserved attached to a mobile host (Vetulicola). The type species is V. gregarius.
WronacyathusGen. et sp. novValidKruse & DebrenneCambrianA member of Archaeocyatha. Genus includes new species W. ayuzhui.
Zhongpingscolex[204] Gen. et sp. novValidShao et al.Cambrian (Fortunian)KuanchuanpuA scalidophoran, probably a stem-group kinorhynch. Genus includes new species Z. qinensis.
Zuunia[205] Gen. et sp. novYang et al.Late EdiacaranZuun-ArtsA cloudinid. The type species is Z. chimidtsereni.

Research

Other organisms

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images
Annularidens[221] Gen. et sp. novIn pressOuyang et al.EdiacaranDoushantuoAn acritarch. Genus includes new species A. inconditus.
Anqiutrichoides[222] Gen. et sp. novValidLi et al.TonianShiwangzhuangA multicellular organism of uncertain phylogenetic placement, possibly a eukaryotic alga. Genus includes new species A. constrictus.
Aphralysia anfracta[223] Sp. novValidKopaska-Merkel, Haywick & KeyesCarboniferous (Serpukhovian)
A tubular calcitic microfossil of uncertain affinities
Arborea denticulata[224] Sp. novValidWang et al.EdiacaranDengyingA frondose fossil of uncertain affinities.
Archaeosporites[225] Gen. et sp. novValidHarper et al.Early DevonianRhynie chertA fungus belonging to the group Archaeosporaceae. Genus includes new species A. rhyniensis.
Asteridium tubulusSp. novValidYin et al.Cambrian Stage 4An organic-walled microfossil. Announced in 2020; the final version of the article naming it was published in 2021.
Attenborites[226] Gen. et sp. novValidDroser et al.EdiacaranRawnsleyAn organism of uncertain phylogenetic placement, described on the basis of a well-defined irregular oval to circular fossil. Genus includes new species A. janeae. Announced in 2018; the final version of the article naming it was published in 2020.
Bispinosphaera vacuaSp. novIn pressOuyang et al.EdiacaranDoushantuoAn acritarch.
Brijax[227] Gen. et sp. novIn pressKrings & HarperDevonianRhynie chertA probable chytrid fungus. Genus includes new species B. amictus.
Convolutubus[228] Gen. et sp. novValidVaziri et al.EdiacaranAn organic-walled tubular organism. Genus includes new species C. dargazinensis.
Corrugasphaera perfectaSp. novValidYin et al.Cambrian Stage 4An organic-walled microfossil. Announced in 2020; the final version of the article naming it was published in 2021.
CrassimembranaGen. et 2 sp. novIn pressOuyang et al.EdiacaranDoushantuoAn acritarch. Genus includes new species C. crispans and C. multitunica.
Cyanosarcinopsis[229] Gen. et sp. novValidCalça & FairchildPermianAssistênciaA chroococcacean. Genus includes new species C. hachiroi.
Cyathochitina brussai[230] Sp. novIn pressDe la Puente, Paris & VaccariOrdovician (Hirnantian) and Silurian (Rhuddanian)Brutia
Clemville
Salar del Rincón
Soom Shale





?
?
?		A chitinozoan.
Cyathochitina lariensisSp. novIn pressDe la Puente, Paris & VaccariLatest Ordovician–earliest SilurianSalar del RincónA chitinozoan.
Cyathochitina punaensisSp. novIn pressDe la Puente, Paris & VaccariLatest Ordovician–earliest SilurianSalar del RincónA chitinozoan.
Cymatiosphaera spinaSp. novValidYin et al.Cambrian Stage 4An organic-walled microfossil. Announced in 2020; the final version of the article naming it was published in 2021.
Dichothallus[231] Gen. et sp. novIn pressNaugolnykhPermian (early Kungurian)PhilippovianA brown alga of uncertain phylogenetic placement. Genus includes new species D. divaricatus.
Dictyocyrillium[232] Gen. et sp. novIn pressMartí Mus, Moczydłowska & KnollTonianElbobreenA vase-shaped microfossil. Genus includes new species D. erythron.
Distosphaera jinguadunensisSp. novIn pressOuyang et al.EdiacaranDoushantuoAn acritarch.
Dongyesphaera[233] Gen. et sp. novIn pressYin et al.PaleoproterozoicTianpengnaoAn acritarch. Genus includes new species D. tenuispina.
Eoentophysalis hutuoensisSp. novIn pressYin et al.PaleoproterozoicHebiancunA cyanobacterium belonging to the family Entophysalidaceae
Eosolena magnaSp. novValidLi et al.TonianShiwangzhuangA multicellular, eukaryotic alga.
Flabellophyton obesum[234] Sp. novValidWan et al.EdiacaranAn organism of uncertain phylogenetic placement, possibly an alga.
Flabellophyton stupendum[235] Sp. novIn pressXiao et al.EdiacaranRawnsley QuartziteProbably a benthic macroalga.
Flabellophyton typicumSp. novValidWan et al.EdiacaranAn organism of uncertain phylogenetic placement, possibly an alga.
Liulingjitaenia irregularisSp. novIn pressXiao et al.EdiacaranRawnsley QuartziteProbably a benthic macroalga.
Mengeosphaera matryoshkaformisSp. novIn pressOuyang et al.EdiacaranDoushantuoAn acritarch.
Nepia[236] Gen. et sp. novValidGolubkova in Golubkova & KochnevEdiacaranAn oscillatorian cyanobacteria. Genus includes new species N. calicina.
Noffkarkys[237] Gen. et sp. novValidRetallack & BrozEdiacaran and CambrianArumbera
Flathead
Grant Bluff
Jodhpur
Synalds



An organism of uncertain phylogenetic placement, a member of the family Charniidae. Genus includes new species N. storaaslii. Announced in 2020; the final version of the article naming it was published in 2021.
Obamus[238] Gen. et sp. novValidDzaugis et al.EdiacaranRawnsleyA torus-shaped organism, similar in gross morphology to some poriferans and benthic cnidarians. Genus includes new species O. coronatus. Announced in 2018; the final version of the article naming it was published in 2020.
Ophiocordyceps dominicanus[239] Sp. novValidPoinar & VegaEocene or MioceneDominican amberA fungus, a species of Ophiocordyceps. Announced in 2019; the final version of the article naming it was published in 2020.
Palaeomycus[240] Gen. et sp. novValidPoinarLate Cretaceous (Cenomanian)Burmese amberA fungus described on the basis of pycnidia. Genus includes new species P. epallelus. Announced in 2018; the final version of the article naming it was published in 2020.
Pararenicola gejiazhuangensisSp. novValidLi et al.TonianShiwangzhuangA coenocytic alga.
Patagonifilum[241] Gen. et sp. novIn pressMassini et al.Late JurassicLa MatildeA cyanobacterium. Genus includes new species P. jurassicum.
Plagasphaera[242] Gen. et sp. novValidYin et al.Cambrian Stage 4An organic-walled microfossil. Genus includes new species P. balangensis. Announced in 2020; the final version of the article naming it was published in 2021.
Polycephalomyces balticaSp. novValidPoinar & VegaEoceneBaltic amber
A fungus belonging to the family Ophiocordycipitaceae. Announced in 2019; the final version of the article naming it was published in 2020.
Proaulopora ordosia[243] Sp. novIn pressLiu et al.OrdovicianOrdos BasinA member of Nostocales.
Protoarenicola baishicunensisSp. novValidLi et al.TonianShiwangzhuangA coenocytic alga.
Protoarenicola shijiacunensisSp. novValidLi et al.TonianShiwangzhuangA coenocytic alga.
Protographum[244] Gen. et sp. novValidLe Renard et al.Early CretaceousPotomac
A fungus belonging or related to the family Aulographaceae. Genus includes new species P. luttrellii.
Pterospermella vinctusaSp. novValidYin et al.Cambrian Stage 4An organic-walled microfossil. Announced in 2020; the final version of the article naming it was published in 2021.
Ramochitina deynouxiSp. novIn pressDe la Puente, Paris & VaccariLatest Ordovician–earliest SilurianSalar del Rincón
A chitinozoan.
Sinosabellidites huangshanensisSp. novValidLi et al.TonianShiwangzhuangA coenocytic alga.
Spinachitina titaeSp. novIn pressDe la Puente, Paris & VaccariLatest Ordovician–earliest SilurianSalar del RincónA chitinozoan.
Spiroplasma burmanica[245] Gen. et sp. novValidPoinarCretaceous (Albian-Cenomanian)Burmese amberA bacterium belonging to the group Mollicutes, a species of Spiroplasma.
Stomiopeltites shangcunicus[246] Sp. novValidMaslova & Tobias in Maslova et al.OligoceneShangcunA fungus belonging to the family Micropeltidaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Triskelia[247] Gen. et sp. novValidStrullu-Derrien et al.DevonianRhynie ChertAn organism of uncertain phylogenetic placement, possibly a green alga or a fungus.[248] Genus includes new species T. scotlandica. Announced in 2020; the final version of the article naming it was published in 2021.
Windipila wimmervoecksii[249] Sp. novValidKrings & HarperEarly DevonianWindyfieldA fungal reproductive unit. Announced in 2019; the final version of the article naming it was published in 2020.

Research

Trace fossils

History of life in general

Other research

Notes and References

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  60. José Amet Rivaz Hernández . 2020 . Linnaeocaninella nomen novum for the Middle Permian fossil Caninella Liang, 1990 (Brachiopoda: Productida: Richthofenidae), preoccupied by Caninella Gorsky, 1938 (Cnidaria: Anthozoa: Bothrophyllidae) . Zootaxa . 4732 . 2 . 335–336 . 10.11646/zootaxa.4732.2.9 . 32230266 . 214382236 .
  61. Xiaolin Duan . Yue Liang . Lars E. Holmer . Zhifei Zhang . 2020 . First report of acrotretoid brachiopod shell beds in the lower Cambrian (Stage 4) Guanshan Biota of eastern Yunnan, South China . Journal of Paleontology . 95 . 1 . 40–55 . 10.1017/jpa.2020.66 . 225289176 .
  62. V. V. Baranov . 2020 . New rhynchonellids and atrypids (Brachiopoda) from the Lower Devonian deposits of Northeast Eurasia . Paleontological Journal . 54 . 3 . 263–272 . 10.1134/S003103012003003X . 219958612 .
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  64. Huiting Wu . Yang Zhang . Thomas L. Stubbs . Jingqi Liu . Yuanlin Sun . 2020 . A new Changhsingian (Lopingian) brachiopod fauna of the shallow-water clastic shelf facies from Fujian Province, south-eastern China . Papers in Palaeontology . 7 . 2 . 861–884 . 10.1002/spp2.1318 . 225720486 .
  65. Luiz Felipe Aquino Corrêa . Maria Inês Feijó Ramos . 2020 . Discinoids (Brachiopoda: Lingulata) from the upper Manacapuru Formation (Early Devonian), south border of Amazonas Basin, Brazil . Journal of South American Earth Sciences . 105 . Article 102960 . 10.1016/j.jsames.2020.102960 . 226351592 .
  66. Zhiliang Zhang . Lars E. Holmer . Feiyang Chen . Glenn A. Brock . 2020 . Ontogeny and evolutionary significance of a new acrotretide brachiopod genus from Cambrian Series 2 of South China . Journal of Systematic Palaeontology . 18 . 19 . 1569–1588 . 10.1080/14772019.2020.1794991 . 225469021 . free .
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