2020 in paleoichthyology explained

This list of fossil fishes described in 2020 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2020, as well as other significant discoveries and events related to paleoichthyology that occurred in 2020.

New taxa

Jawless vertebrates

Name	Novelty	Status	Authors	Age	Type locality	Notes
Clavulaspis^[1]	Nom. nov	Valid	Elliott, Lassiter & Geyer	Devonian (Givetian)	Yahatinda Formation	A member of the family Pteraspididae; a replacement name for Helaspis Elliott et al. (2000).
Ecphymaspis	Gen. et comb. nov	Valid	Elliott, Lassiter & Geyer	Devonian (Eifelian)	Spring Mountain beds	A member of the family Pteraspididae; a new genus for "Psephaspis" idahoensis Denison (1968).
Parathelodus liaokuoensis^[2]	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xishancun Xitun	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Parathelodus wangi	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xishancun Xitun	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Parathelodus xitunensis	Sp. nov	Valid	Cui et al.	Devonian (Lochkovian)	Xitun	A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
Psarkosteus^[3]	Gen. et sp. nov	Valid	Dec	Devonian (Givetian)	Skały	A member of the family Psammosteidae. The type species is P. mediocris.
Rumporostralis^[4]	Gen. et comb. et sp. nov	Valid	Shan et al.	Silurian (Telychian)	Xikeng Formation	A member of Galeaspida belonging to the group Eugaleaspiformes and the family Sinogaleaspidae. The type species is "Sinogaleaspis" xikengensis Pan & Wang (1980); genus also includes new species R. shipanensis.
Scutellaspis	Gen. et sp. nov	Valid	Elliott, Lassiter & Geyer	Devonian (Eifelian)	Spring Mountain beds	A member of the family Pteraspididae. Genus includes new species S. wilsoni.

Placoderms

Name	Novelty	Status	Authors	Age	Type locality	Notes
Actinolepis zaikai^[5]	Sp. nov	Valid	Plax & Newman	Devonian (Emsian)	Lepel Beds
Bulongosteus^[6]	Gen. et sp. nov	Valid	Liu, Zong & Gong	Late Devonian	Zhulumute Formation	A member of the family Selenosteidae. Genus includes new species B. liui.
Johannaspis^[7]	Gen. et comb. nov	Valid	Vaškaninová	Early Devonian	Prague Basin	A homostiid arthrodire. Genus includes "Asterolepis" bohemicus Barrande (1872).
Minjinia^[8]	Gen. et sp. nov	Valid	Brazeau et al.	Devonian (Pragian)		A placoderm closely related to the gnathostome crown group. Genus includes new species M. turgenensis.
Stipatosteus	Gen. et sp. nov	Valid	Plax & Newman	Devonian (Emsian)	Lepel Beds	A phlyctaeniid arthrodire. Genus includes new species S. svidunovitchi.

Cartilaginous fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Amamriabatis^[10]	Gen. et sp. nov	Valid	Adnet et al.	Eocene		?	A ray belonging to the family Mobulidae. The type species is A. heni.
Asteracanthus dunaii^[11]	Sp. nov	Valid	Szabó & Főzy	Jurassic
Atlantitrygon^[12]	Gen. et comb. et sp. nov	Valid	Sambou et al.	Paleocene (Thanetian) and Eocene (Lutetian)	Matam		A putative marine potamotrygonid. The type species is "Dasyatis" sudrei Cappetta (1972); genus also includes new species A. senegalensis.
Canadodus^[13]	Gen. et sp. nov	Valid	Popov, Johns & Suntok	Late Oligocene	Sooke		A member of the family Chimaeridae. Genus includes new species C. suntoki.
Carcharhinus kasserinensis	Sp. nov	Valid	Adnet et al.	Eocene (Bartonian)		?	A species of Carcharhinus.
Coupatezia cristata	Sp. nov	Valid	Adnet et al.	Eocene			A ray belonging to the superfamily Dasyatoidea.
Cretasquatina^[14]	Gen. et sp. nov	Valid	Maisey, Ehret & Denton	Late Cretaceous (Campanian)	Mooreville Chalk Navesink Formation?	( ?)	A member of the family Squatinidae. The type species is C. americana.
Eoplatyrhina^[15]	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca		A member of the family Platyrhinidae. Genus includes "Platyrhina" bolcensis Heckel (1851).
Eorhinobatos^[16]	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca		A guitarfish; a new genus for "Rhinobatos" primaevus.
Ferromirum^[17]	Gen. et sp. nov	Valid	Frey et al.	Devonian (Famennian)	Ibâouane		A member of Symmoriiformes. The type species is F. oukherbouchi.
Himantura souarfortuna	Sp. nov	Valid	Adnet et al.	Eocene (Bartonian)			A whiptail stingray, a species of Himantura.
Lamarodus^[18]	Gen. et sp. nov	Valid	Ivanov in Ivanov et al.	Permian (Guadalupian)	Bell Canyon Formation		A member of Hybodontiformes belonging to the superfamily Hybodontoidea. Genus includes new species L. triangulus. Announced in 2018; the final version of the article naming it was published in 2020.
Leptocharias tunisiensis	Sp. nov	Valid	Adnet et al.	Eocene		? ?	A relative of the barbeled houndshark.
Mecotrygon	Gen. et sp. nov	Valid	Adnet et al.	Eocene			A whiptail stingray belonging to the subfamily Neotrygoninae. The type species is M. asperodentulus.
Mennerotodus mackayi^[19]	Sp. nov	Valid	Cicimurri, Ebersole & Martin	Paleocene (Danian)	Clayton		A sand shark
Mennerotodus parmleyi	Sp. nov	Valid	Cicimurri, Ebersole & Martin	Eocene (Bartonian)	Clinchfield		A sand shark
Microtriftis	Gen. et sp. nov	Valid	Sambou et al.	Eocene (Ypresian to Lutetian)	Matam		A dasyatoid of uncertain phylogenetic placement. Genus includes new species M. matami.
Nanocetorhinus zeitlingeri^[20]	Sp. nov	Valid	Feichtinger, Pollerspöck & Harzhauser	Oligocene (Chattian)	Eferding		A member of Neoselachii of uncertain phylogenetic placement.
Odontorhytis priemi	Sp. nov	Valid	Sambou et al.	Paleocene (Thanetian) and Eocene (Ypresian)	Matam		A member of Neoselachii of uncertain phylogenetic placement
Ouledia lacuna	Sp. nov	Valid	Adnet et al.	Eocene (Bartonian)			A butterfly ray.
Pachygymnura	Gen. et comb. nov	Valid	Adnet et al.	Eocene			A butterfly ray. The type species is "Coupatezia" attiai Cook in Murray et al. (2010).
Plesiozanobatus	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca		A relative of panray. Genus includes "Torpedo" egertoni De Zigno (1876).
Pristiophorus austriacus^[21]	Sp. nov	Valid	Reinecke et al.	Miocene (Aquitanian)	Ebelsberg		A species of Pristiophorus.
Pristiophorus borealis	Sp. nov	Valid	Reinecke et al.	Oligocene (Chattian)	Sülstorf		A species of Pristiophorus.
Pristiophorus tortonicus	Sp. nov	Valid	Reinecke et al.	Miocene (Tortonian)	Mica Clay		A species of Pristiophorus.
Pristiophorus ungeri	Sp. nov	Valid	Reinecke et al.	Miocene (Burdigalian)	Neuhofen		A species of Pristiophorus.
Pseudorhinobatos	Gen. et comb. nov	Valid	Marramà et al.	Eocene	Monte Bolca		A guitarfish; a new genus for "Rhinobatos" dezignii.
Sculptospina^[22]	Gen. et sp. nov	Valid	Lebedev in Lebedev, Ivanov & Linkevich	Devonian (Famennian)			A member of Ctenacanthiformes of uncertain phylogenetic placement. The type species is S. makhlaevi.
Sphyrna guinoti	Sp. nov	Valid	Adnet et al.	Eocene		?	A species of Sphyrna.
Stegostoma tethysiensis	Sp. nov	Valid	Adnet et al.	Eocene		?	A relative of the zebra shark.
Taklamakanolepis^[23]	Gen. et sp. nov		Andreev et al.	Early Silurian	Ymogantau		A member of Mongolepidida. The type species is T. asiaticus.
Tielikewatielepis	Gen. et sp. nov		Andreev et al.	Early Silurian	Tataertag Ymogantau		A member of Mongolepidida. The type species is T. sinensis.
Xiaohaizilepis	Gen. et sp. nov		Andreev et al.	Early Silurian	Tataertag Ymogantau		A member of Mongolepidida. The type species is X. liui.
Yuanolepis	Gen. et sp. nov		Andreev et al.	Early Silurian	Ymogantau		A cartilaginous fish of uncertain phylogenetic placement. The type species is Y. bachunensis.

Ray-finned fishes

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Agassizilia^[24]	Gen. et sp. nov	Valid	Cooper & Martill	Cenomanian	Kem Kem		Possibly a member of the family Pycnodontidae. The type species is A. erfoudina.
Altmuehlfuro^[25]	Gen. et sp. nov	Valid	Ebert	Late Jurassic	Solnhofen		A member of Halecomorphi belonging to the group Ophiopsiformes. Genus includes new species A. boomerang.
Amakusaichthys^[26]	Gen. et sp. nov	Valid	Yabumoto, Hirose & Brito	Late Cretaceous (Santonian)	Hinoshima		A member of Ichthyodectiformes. Genus includes new species A. goshouraensis. Announced in 2018; the final version of the article naming it was published in 2020.
Ampheristus americanus	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian) and Paleocene (Danian)	Clayton Kemp Clay	( )	A cusk-eel.
Anguilla? chickasawae^[27]	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian) and Paleocene (Danian)	Clayton Ripley	( )	A member or a relative of the family Anguillidae
Anomoeodus wolfi^[28]	Sp. nov	Valid	Capasso	Late Cretaceous (Cenomanian)	Del Rio		A member of the family Pycnodontidae.
Anorevus^[29]	Gen. et sp. nov.	Valid	Bannikov & Zorzin	Eocene (Ypresian)	Monte Bolca		A member of Percomorpha of uncertain phylogenetic placement. The type species is A. lorenzonii.
Apateodus crenellatus	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley
Ariopsis castilloensi^[30]	Sp. nov	In press	Aguilera & Rodriguez in Aguilera et al.	Late Oligocene to early Miocene	Castillo		A species of Ariopsis.
Armigatus carrenoae^[31]	Sp. nov	Valid	Alvarado-Ortega, Than-Marchese & Melgarejo-Damián	Early Cretaceous (Albian)	Tlayúa		A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Armigatidae.
Bagre ornatus	Sp. nov	In press	Aguilera & Rodriguez in Aguilera et al.	Late Oligocene to early Miocene	Castillo		A species of Bagre.
Baringochromis^[32]	Gen. et 3 sp. nov	Valid	Altner & Reichenbacher	Late Miocene			A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is B. senutae; genus also includes B. sonyii and B. tallamae. Announced in 2020; the final version of the article naming it was published in 2021.
Benthophilus aprutinus	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A tadpole goby.
Benthophilus labronicus	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A tadpole goby.
Boreolates^[33]	Gen. et sp. nov	Valid	Weems	Eocene	Nanjemoy		A member of the family Latidae. The type species is B. debernardi.
Brachyplatystoma elbakyani^[34]	Sp. nov	In press	Agnolin & Bogan	Late Miocene			A species of Brachyplatystoma
Buenia pisiformis	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian and Messinian)			A species of Buenia.
Burguklia minichorum^[35]	Sp. nov	Valid	Bakaev & Kogan	Permian (Wordian–Capitanian)			An early ray-finned fish. Announced in 2019; the final version of the article naming it was published in 2020.
Caspiosoma lini	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A species of Caspiosoma.
Caspiosoma paulisulcata	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A species of Caspiosoma.
Caucombrus^[36]	Gen. et sp. nov	Valid	Bannikov	Early Oligocene		Abkhazia	A member of the family Scombridae. Genus includes new species C. histiopterygius.
Chaychanus^[37]	Gen. et sp. nov	Valid	Cantalice Severiano, Alvarado Ortega & Bellwood	Paleocene			A member of the family Pomacentridae. Genus includes new species C. gonzalezorum. Announced in 2019; the final version of the article naming it was published in 2020.
Cheirolepis aleshkai^[38]	Sp. nov	Valid	Plax	Devonian (Eifelian)			An early ray-finned fish.
Choctawichthys	Gen. et comb. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley		A member of Paracanthopterygii of uncertain phylogenetic placement. The type species is "genus Perciformorum" cepoloides Nolf & Dockery (1990).
Chromogobius? primigenius^[39]	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene			Possibly a species of Chromogobius.
Clupea hanishinaensis^[40]	Nom. nov	Valid	Yabumoto & Nazarkin	Miocene	Bessho		A species of Clupea; a replacement name for Clupea macrocephala Yabumoto & Nazarkin (2018).
Congrophichthus	Gen. et sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (late Campanian and Maastrichtian) and Paleocene (Danian)	Clayton Coffee Sand Kemp Clay	( )	A member of the family Congridae. The type species is C. transterminus.
Cowetaichthys carnevalei	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley		A beardfish
Cyclothone duhoensis^[41]	Sp. nov	Valid	Nam & Nazarkin	Miocene	Duho		A species of Cyclothone. Announced in 2020; the final version of the article naming it was published in 2021.
Dakotaichthys	Gen. et sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian)	Fox Hills Kemp Clay	( )	Possibly a member of the family Gadidae. The type species is D. hogansoni.
Eleotris omuamuaensis^[42]	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian)			A species of Eleotris.
Eleotris tyrrhenicus	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian)			A species of Eleotris.
Ellimma longipectoralis^[43]	Sp. nov	Valid	Polck et al.	Early Cretaceous (Aptian)	Barra Velha Formation		A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Announced in 2019; the final version of the article naming it was published in 2020.
Ellimmichthys spinosus^[44]	Sp. nov	In press	De Figueiredo & Gallo	Early Cretaceous	Recôncavo Basin		A member of Clupeomorpha belonging to the group Ellimmichthyiformes.
Elongofuro	Gen. et sp. nov	Valid	Ebert	Late Jurassic			A member of Halecomorphi belonging to the group Ophiopsiformes. Genus includes new species E. woelflei.
Elopothrissus carsonsloani^[45]	Sp. nov	Valid	Schwarzhans & Stringer	Paleocene (Danian)	Clayton		A relative of the Japanese gissu.
Engdahlichthys^[46]	Gen. et sp. nov	Valid	Murray et al.	Paleocene	Fort Union		A sturgeon. Genus includes new species E. milviaegis.
Enigmacottus	Gen. et sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			Possibly a member of the family Psychrolutidae. The type species is E. socialis.
Epaelops^[47]	Gen. et sp. nov	In press	Alves, Alvarado Ortega & Brito	Early Cretaceous (Albian)	Tlayúa		A member of Elopiformes. Genus includes new species E. martinezi. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Eutawichthys choctawae	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Kemp Clay Ripley	( )	Probably a member of Beryciformes
Feroxichthys^[48]	Gen. et sp. nov	Valid	Xu	Middle Triassic (Anisian)	Guanling		A member of the family Colobodontidae. The type species is F. yunnanensis.
Globanomoeodus	Gen. et sp. nov	Valid	Capasso	Late Cretaceous (Cenomanian)	Del Rio		A member of the family Pycnodontidae. The type species is G. dentespassim.
Gobius peloponnesus	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Pliocene (Zanclean)			A species of Gobius.
Gobius supraspectabilis	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene			A species of Gobius.
Gregoriopycnodus^[49]	Gen. et comb. nov	Valid	Taverne, Capasso & Del Re	Early Cretaceous (Albian)			A member of the family Pycnodontidae. The type species is "Palaeobalistum" bassanii d'Erasmo (1914).
Hadromos^[50]	Gen. et sp. nov	Valid	Murray	Early Cenozoic (possibly Eocene)	Sangkarewang		A member of Cyprinoidea. Genus includes new species H. sandersae.
Hastichthys totonacus^[51]	Sp. nov	In press	Alvarado-Ortega & Díaz-Cruz	Late Cretaceous (Turonian)	Agua Nueva		A member of Aulopiformes belonging to the family Dercetidae.
Heckelichthys preopercularis^[52]	Sp. nov	In press	Baños Rodríguez et al.	Cretaceous (Albian–Cenomanian)	El Doctor		A member of Ichthyodectiformes
Hesperichthys gironeae	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian and Messinian)			A goby.
Hesperichthys iugosus	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene			A goby. Originally described as a species of Hesperichthys, but subsequently transferred to the genus Sarmatigobius.^[53]
Hiascoactinus^[54]	Gen. et sp. nov	In press	Kim et al.	Late Triassic	Amisan		A member of Redfieldiiformes. Genus includes new species H. boryeongensis. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Hoeseichthys	Gen. et comb. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene and Pliocene		^[55] ^[56]	A goby. Genus includes "Otolithus (Gobius)" praeclarus Procházka (1893) and H. laevis (Weiler, 1942), as well as "Gobiida" bicornuta Lin, Girone & Wolf (2015) and "Gobiida" brioche Lin, Girone & Wolf (2015).
Holzmadenfuro^[57]	Gen. et sp. nov	Valid	Ebert, Thies & Hauff	Early Jurassic (Toarcian)	Posidonia		A member of Halecomorphi. Genus includes new species H. rebmanni.
Hoplopteryx langfordi	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley
Hypsocormus posterodorsalis^[58]	Sp. nov	Valid	Maxwell et al.	Late Jurassic (Kimmeridgian)	Nusplingen
Knipowitschia etrusca	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A species of Knipowitschia.
Knipowitschia polonica	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene			A species of Knipowitschia.
Kokenichthys navis	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley		A member of Osteoglossiformes of uncertain phylogenetic placement
Kradimus^[59]	Gen. et sp. nov	Valid	Veysey, Brito & Martill	Late Cretaceous (Turonian)	Akrabou Formation		A member of Crossognathiformes. Genus includes new species K. asflaensis. Announced in 2019; the final version of the article naming it was published in 2020.
Lavinia stuwilliamsi^[60]	Sp. nov	Valid	McClellan & Smith	Late Miocene	Salt Lake Formation		A species of Lavinia.
Lebrunichthys^[61]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)			A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is L. nammourensis.
Lesueurigobius stazzanensis	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Tortonian)			A species of Lesueurigobius.
Lindoeichthys^[62]	Gen. et sp. nov.	Valid	Murray et al.	Late Cretaceous (Maastrichtian)	Scollard Formation		A member of Percopsiformes. Genus includes new species L. albertensis. Announced in 2019; the final version of the article naming it was published in 2020.
Louwoichthys^[63]	Gen. et sp. nov	Valid	Xu	Middle Triassic (Anisian)			A member of the stem group of Neopterygii. Genus includes new species L. pusillus.
Luganoia fortuna^[64]	Sp. nov	Valid	Xu	Middle Triassic (Ladinian)	Falang
Megalops? nolfi	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley		Probably member of the family Megalopidae
Monosmilus^[65]	Gen. et sp. nov		Capobianco et al.	Eocene (Lutetian)	Domanda		A stem-engraulid. The type species is M. chureloides.
Muraenanguilla unionensis	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley		An eel of uncertain phylogenetic placement
Neilpeartia^[66]	Gen. et sp. nov	Valid	Carnevale et al.	Eocene (Ypresian)	Monte Bolca		A frogfish. Genus includes new species N. ceratoi.
Neomesturus^[67]	Gen. et sp. nov	In press	Cooper & Martill	Turonian	Akrabou		A member of Pycnodontiformes. Genus includes new species N. asflaensis.
Neoproscinetes africanus	Sp. nov	Valid	Cooper & Martill	Cenomanian	Kem Kem		A member of the family Pycnodontidae
Njoerdichthys^[68]	Gen. et sp. nov	In press	Cawley et al.	Late Cretaceous (Turonian)	Hesseltal		A member of the family Pycnodontidae. The type species is N. dyckerhoffi.
Nunaneichthys^[69]	Gen. et sp. nov	In press	Hernández-Guerrero et al.	Cretaceous (Albian-Cenomanian)			A bonefish belonging to the subfamily Pterothrissinae. Genus includes new species N. mexicanus.
Odondebuenia agiadiae	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene			A relative of the coralline goby.
Ohmdenfuro	Gen. et sp. nov	Valid	Ebert, Thies & Hauff	Early Jurassic (Toarcian)	Posidonia		A member of Halecomorphi. Genus includes new species O. bodmani.
Osmeroides mississippiensis	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley
Ossulcus	Gen. et sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley		Probably a member of Beryciformes. The type species is O. labiatus.
Otolithopsis cumatilis	Sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley		An otolith of a fish of uncertain phylogenetic placement.
Palaeogadus weltoni	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian)	Kemp Clay
Palealbula korchinskyi^[70]	Sp. nov	Valid	Schwarzhans & Mironenko	Late Jurassic			Possibly a stem-albuliform
Palealbula moscoviensis	Sp. nov	Valid	Schwarzhans & Mironenko	Late Jurassic			Possibly a stem-albuliform.
Paranursallia cavini	Sp. nov	In press	Cooper & Martill	Turonian	Akrabou		A member of Pycnodontiformes.
Pauciuncus	Gen. et comb. nov	Valid	Murray	Early Cenozoic (possibly Eocene)	Sangkarewang		A member of Cyprinoidea. Genus includes "Puntius" bussyi.
Polcynichthys^[71]	Gen. et sp. nov	In press	London & Shimada	Late Cretaceous (Cenomanian)	Tarrant		A member of the family Pachyrhizodontidae. Genus includes new species P. lloydhilli.
Proterorhinus cretensis	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Pliocene (Zanclean)			A species of Proterorhinus.
Pythonichthys arkansasensis	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian) and Paleocene (Danian)	Clayton Kemp Clay	( )	A species of Pythonichthys.
Rachycentron stremphaencus^[72]	Sp. nov	Valid	Godfrey & Carnevale	Miocene (Tortonian)	St. Marys		A relative of the cobia.
Rebekkachromis valyricus^[73]	Sp. nov	Valid	Kevrekidis & Reichenbacher in Kevrekidis et al.	Miocene	Ngorora		A cichlid belonging to the tribe Oreochromini.
Rebekkachromis vancouveringae	Sp. nov	Valid	Kevrekidis & Reichenbacher in Kevrekidis et al.	Miocene	Ngorora		A cichlid belonging to the tribe Oreochromini.
Rhynchoconger brettwoodwardi	Sp. nov	Valid	Schwarzhans & Stringer	Late Cretaceous (Maastrichtian)	Kemp Clay		A species of Rhynchoconger.
Rupelia^[74]	Gen. et comb. nov	Valid	Baykina & Kovalchuk in Kovalchuk et al.	Oligocene (Rupelian)			A member of family Clupeidae. Genus includes R. rata (Daniltshenko, 1959).
Salwaichthys^[75]	Gen. et sp. nov	Valid	Bannikov	Early Oligocene		Abkhazia	A member of Perciformes belonging to the new family Salwaichthyidae. Genus includes new species S. paratethyensis.
Sanctusichthys^[76]	Gen. et sp. nov	Valid	López-Arbarello, Maxwell & Schweigert	Late Jurassic (Kimmeridgian)	Nusplingen Limestone		A member of Halecomorphi. Genus includes new species S. rieteri.
Sangkarewangia	Gen. et sp. nov	Valid	Murray	Early Cenozoic (possibly Eocene)	Sangkarewang		A member of Cyprinoidea. Genus includes new species S. sumatranus.
Scleropages sanshuiensis^[77]	Sp. nov	Valid	Zhang	Early Eocene	Huachong		A species of Scleropages. Announced in 2019; the final version of the article naming it was published in 2020.
Scombroclupea javieri^[78]	Sp. nov	In press	Than Marchese et al.	Late Cretaceous (Cenomanian)	Cintalapa		A member of Clupeomorpha of uncertain phylogenetic placement
Siniperca ikikoku^[79]	Sp. nov	Valid	Yabumoto	Miocene			A species of Siniperca.
Simocormus	Gen. et sp. nov	Valid	Maxwell et al.	Late Jurassic (Kimmeridgian–Tithonian)			A member of the family Pachycormidae. The type species is S. macrolepidotus.
Stanhopeichthys^[80]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)			A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is S. libanicus.
Stefanichthys^[81]	Gen. et sp. nov.	Valid	Bannikov & Zorzin	Eocene (Ypresian)	Monte Bolca		A member of Percoidei of uncertain phylogenetic placement. The type species is S. mariannae.
Thorogobius petilus	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A species of Thorogobius.
Thrax	Gen. et sp. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Ripley		A member of Aulopiformes belonging to the family Ichthyotringidae. The type species is T. acutus.
Tippaha	Gen. et sp. et comb. nov	Valid	Schwarzhans & Stringer in Stringer et al.	Late Cretaceous (Maastrichtian)	Owl Creek Ripley		A member of Holocentriformes of uncertain phylogenetic placement. The type species is T. mythica; genus also includes "genus Ophidiidarum" cavatus Nolf & Stringer (1996).
Vachalia^[82]	Gen. et sp. nov	Valid	Přikryl & Carnevale	Early Oligocene			A member of the family Platytroctidae. Genus includes new species V. moraviensis.
Vanderhorstia prochazkai	Sp. nov	Valid	Schwarzhans, Brzobohatý & Radwańska	Miocene		^[83]	A species of Vanderhorstia.
Vegrandichthys^[84]	Gen. et sp. nov	Valid	Díaz-Cruz, Alvarado-Ortega & Giles	Late Cretaceous (Cenomanian)	Cintalapa		A member of Aulopiformes belonging to the family Enchodontidae. The type species is V. coitecus.
Warilochromis^[85]	Gen. et sp. nov	Valid	Altner, Ruthensteiner & Reichenbacher	Late Miocene	Ngorora		A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is W. unicuspidatus.
Zosterisessor exsul	Sp. nov	Valid	Schwarzhans, Agiadi & Carnevale	Miocene (Messinian)			A relative of the grass goby.

Lobe-finned fishes

Name	Novelty	Status	Authors	Age	Type locality	Notes
Ferganoceratodus annekempae^[86]	Sp. nov	Valid	Cavin, Deesri & Chanthasit	Late Jurassic	Phu Kradung	A lungfish.
Megalichthys mullisoni^[87]	Sp. nov	Valid	Downs & Daeschler	Devonian (Famennian)	Catskill
Metaceratodus baibianorum^[88]	Sp. nov	Valid	Panzeri et al.	Late Cretaceous	La Colonia	A lungfish.
Neoceratodus potkooroki^[89]	Sp. nov	Valid	Kemp & Berrell	Cretaceous		A lungfish.
Ptychoceratodus oldhami^[90]	Sp. nov	Valid	Bhat & Ray	Late Triassic (Carnian)	Tiki	A lungfish. Announced in 2018; the final version of the article naming it was published in 2020.
Rossichthys^[91]	Gen. et sp. nov	Valid	Johanson et al.	Carboniferous (Tournaisian)	Ballagan Formation	A member of Rhizodontida. The type species is R. clackae.

Research

A study on the morphology of the osteostracans, evaluating different methods used to determine the morphological variation within this group and its evolution, is published by Ferrón et al. (2020).^[92]
A study on the morphological diversity of osteostracan headshields, aiming to determine the relationship between their morphological diversity and hydrodynamic performance and its implications for the knowledge of the ecological diversity of the osteostracans, is published by Ferrón et al. (2020).^[93]
A study aiming to test the alternative hypotheses of placoderm jaw bone homologies, and evaluating their implications for the knowledge of evolution of jaw bones in early jawed vertebrates, is published by King & Rücklin (2020).^[94]
A study on the fossil dentitions of acanthothoracids is published by Vaškaninová et al. (2020), who report that the teeth of acanthothoracids differed fundamentally from those of arthrodires, and argue that the characteristic traits of acanthothoracid dentition might be ancestral for all jawed vertebrates.^[95]
Redescription of the anatomy of Walterilepis speciosa, based on data from new fossil material, and a study on the phylogenetic relationships of this species is published online by Lukševičs (2020).^[96]
Description of the neurocranial anatomy of Ellopetalichthys scheii is published by Castiello et al. (2020).^[97]
A study aiming to determine whether Titanichthys was a suspension feeder, focusing on mechanical properties of its jaw, is published by Coatham et al. (2020).^[98]
The earliest fossilized vertebrate embryos reported so far, preserved with an adult specimen of Watsonosteus fletti from the Givetian Eday Flagstone Formation (Orcadian Basin; Scotland, United Kingdom), are described by Newman et al. (2020).^[99]
Burrow, Newman & den Blaauwen (2020) describe external spiracular elements in Middle Devonian acanthodians from northern Scotland, differing from spiracles of all known extant and extinct fishes, and report the oldest record of elastic cartilage in the fossil record.^[100]
A study examining the factors influencing the long-term variations of genus-level diversity of elasmobranchs and ray-finned fishes throughout their evolutionary history is published by Guinot & Cavin (2020).^[101]
Carrillo-Briceño et al. (2020) describe a new elasmobranch assemblage from the Oligocene–Miocene boundary in the Dos Bocas Formation (Ecuador), and evaluate the implications of this assemblage for chronostratigraphic inferences and the knowledge of local paleoenvironment.^[102]
Two large vertebrae of sharks belonging to the genus Ptychodus, providing new information on the life history and body size of members of the family Ptychodontidae, are described from the Santonian of Spain by Jambura & Kriwet (2020).^[103]
A study on the evolution of body size in lamniform sharks, including the evolution of gigantism in the lineage of Otodus megalodon, is published by Shimada, Becker & Griffiths (2020).^[104]
Taxonomic revision of the Oligocene and Miocene sand sharks is published by Hovestadt (2020).^[105]
A study aiming to determine the linear body dimensions of Otodus megalodon at different life stages is published by Cooper et al. (2020).^[106]
A study on the class structure of assemblages of specimens of Otodus megalodon in eight previously known formations and in a newly described Miocene locality from northeastern Spain is published by Herraiz et al. (2020), who interpret their findings as indicative of existence of five potential nurseries of these sharks ranging from the Langhian to the Zanclean.^[107]
An isolated tooth of Cosmopolitodus hastalis from the Miocene of South Korea is described by Yun (2020).^[108]
A study on the fossil record of the great white shark from the Pliocene of Peru and Chile is published by Villafaña et al. (2020), who interpret their findings as indicating that great white sharks used the Coquimbo locality in Chile as a nursery and Pisco (Peru) and Caldera (Chile) localities as feeding grounds during the Pliocene.^[109]
A study on the anatomy and phylogenetic relationships of "Urolophus" crassicaudatus is published by Marramà et al. (2020), who transfer this species to the genus Arechia.^[110]
Collareta et al. (2020) describe a fossil stinger a stingray from the Pliocene (Piacenzian) locality La Serra (Italy), twice as long as the longest caudal spines reported from any living stingray species of the Mediterranean Sea, and possibly representing the longest stingray stinger ever reported from both the fossil and the recent records.^[111]
A study on the morphology of the marginal dentition of Lophosteus superbus is published by Chen et al. (2020), who reconstruct the dental ontogeny in this taxon, and evaluate its implications for the knowledge of the evolution of teeth of bony fishes.^[112]
Redescription of the anatomy of Tanyrhinichthys mcallisteri is published by Stack et al. (2020).^[113]
Fragmentary fossil material of Gyrosteus mirabilis is reported from the Toarcian of the Ahrensburg erratics assemblage (Schleswig-Holstein, Germany) by Hornung & Sachs (2020), expanding known geographic range of this species, and representing the first record of a chondrosteid species beyond its type area.^[114]
Redescription of the skeletal anatomy of Yanosteus longidorsalis is published by Hilton, Grande & Jin (2020).^[115]
Revision and a study on the phylogenetic relationships of members of the subfamily Pycnodontinae is published by Poyato-Ariza (2020).^[116]
A study on the skeletal anatomy and phylogenetic relationships of Lombardina decorata is published by Taverne (2020).^[117]
A study on the degree of preservation of the skin of an aspidorhynchid specimen from the Barremian Paja Formation (Colombia), representing the first instance of soft tissue preservation in vertebrates from the Early Cretaceous of northern South America, is published by Alfonso-Rojas & Cadena (2020).^[118]
A study on the diversity and distribution of non-marine teleost fishes in the Western Interior of North America during the late Maastrichtian, based on fossils from the Hell Creek Formation, the Lance Formation and the Scollard Formation, is published online by Brinkman et al. (2020).^[119]
Description of new fossil material of Abisaadichthys libanicus and Eusebichthys byblosi, providing new information on the skeletal anatomy of these taxa, is published by Taverne & Capasso (2020).^[120]
Fossil material of Xiphactinus is described from the latest Maastrichtian Salamanca Formation (Chubut Province, Argentina) by De Pasqua, Agnolin & Bogan (2020), representing the first record of this genus from southern part of South America.^[121]
A methodology for assessing locomotion energetics in extinct bony fishes is presented by Ferrón (2020), who interprets his findings as providing evidence of endothermy in Xiphactinus audax.^[122]
Redescription and a study on the phylogenetic relationships of Laeliichthys ancestralis is published by Brito, Figueiredo & Leal (2020).^[123]
A study on the skeletal anatomy of Pirskenius, aiming to resolve whether Pirskeniidae can be sustained as a separate family, is published by Reichenbacher et al. (2020).^[124]
A study aiming to infer the genetic basis of the reduction of pelvic skeleton in a Miocene stickleback fish Gasterosteus doryssus is published by Stuart, Travis & Bell (2020).^[125]
A fish larva sharing anatomical similarities with the so‐called tholichthys larval stage of butterflyfishes is described from the Eocene (Bartonian) locality of Gornyi Luch (Krasnodar Krai, Russia) by Carnevale & Bannikov (2020).^[126]
New fossil material of Mawsonia gigas, including one of the anatomically most informative specimens referable to the genus Mawsonia, is described from the Mesozoic Tacuarembó Formation (Uruguay) by Toriño et al. (2020).^[127]
New fossil material of Axelrodichthys megadromos is described from several Campanian and Maastrichtian sites in southern France by Cavin et al. (2020), who present a reconstruction of the skull of this species, and study its phylogenetic relationships and ecology.^[128]
Redescription of the anatomy of the skull of Durialepis edentatus is published by Mondéjar‐Fernández, Friedman & Giles (2020).^[129]
Description of new material of tristichopterids from the Devonian (Famennian) locality of Strud (Belgium), and a study on the phylogenetic relationships of tristichopterids, is published by Olive et al. (2020).^[130]
Description of a new, 1.57-metre-long articulated specimen of Elpistostege watsoni from the Upper Devonian of Canada, and a study on the implications of this specimen for the knowledge of the early evolution of the vertebrate hand, is published by Cloutier et al. (2020).^[131]
A study aiming to determine the potential significance of tides for the evolution of bony fish and early tetrapods from the Late Silurian to early Late Devonian is published by Byrne et al. (2020).^[132]
Evidence of enhanced fish production during the extreme global warmth of the early Paleogene is presented by Britten & Sibert (2020).^[133]
A study aiming to determine the impact of changes in the Earth system during the Eocene–Oligocene transition on pelagic fish production and biodiversity is published by Sibert et al. (2020).^[134]

Notes and References

David K. Elliott . Linda S. Lassiter . Kathryn E. Geyer . 2020 . The last pteraspids (Vertebrata, Heterostraci): new material from the Middle Devonian of Alberta and Idaho . Journal of Paleontology . 94 . 4 . 758–772 . 10.1017/jpa.2020.9 . 2020JPal...94..758E . 218776355 .
Xin-Dong Cui . Qiang Li . Tuo Qiao . Min Zhu . 2020 . New material of thelodonts from Lochkovian (Lower Devonian) of Qujing, Yunnan, China . Vertebrata PalAsiatica . 58 . 1 . 1–15 . 10.19615/j.cnki.1000-3118.190612 .
Marek Dec . 2020 . A new Middle Devonian (Givetian) psammosteid (Vertebrata: Heterostraci) from Poland . Annales Societatis Geologorum Poloniae . 90 . 1 . 75–93 . 10.14241/asgp.2020.04 . 225233821 . free .
Xianren Shan . Min Zhu . Wenjin Zhao . Zhaohui Pan . Pingli Wang . Zhikun Gai . 2020 . A new genus of sinogaleaspids (Galeaspida, stem-Gnathostomata) from the Silurian Period in Jiangxi, China . PeerJ . 8 . e9008 . 10.7717/peerj.9008 . 32461826 . 7233274 . free .
Dmitry P. Plax . Michael J. Newman . 2020 . New Early Devonian (late Emsian) placoderms from Belarus . Journal of Paleontology . 94 . 4 . 773–787 . 10.1017/jpa.2020.6 . 2020JPal...94..773P . 216191250 .
Yilong Liu . Ruiwen Zong . Yiming Gong . 2020 . The first aspinothoracid arthrodire from the Late Devonian continental strata are found in western Junggar, Xinjiang . Chinese Science Bulletin . 65 . 21 . 2256–2265 . 10.1360/TB-2020-0108 . 225991890 . free .
Valéria Vaškaninová . 2020 . New genus of homostiid arthrodire contributes to the revision of placoderm diversity in the Early Devonian of the Prague Basin . Alcheringa: An Australasian Journal of Palaeontology . 44 . 3 . 397–410 . 10.1080/03115518.2020.1749303 . 2020Alch...44..397V . 219456262 .
Martin D. Brazeau . Sam Giles . Richard P. Dearden . Anna Jerve . Ya. Ariunchimeg . E. Zorig . Robert Sansom . Thomas Guillerme . Marco Castiello . 2020 . Endochondral bone in an Early Devonian 'placoderm' from Mongolia . Nature Ecology & Evolution . 4 . 11 . 1477–1484 . 10.1038/s41559-020-01290-2 . 32895518 . 2020NatEE...4.1477B . 221543148 . 10044/1/82739 . free .
Michael J. Newman . Jan L. Den Blaauwen . Carole J. Burrow . 2020 . Two newly identified cheiracanthid acanthodians from the Mey Flagstone Formation (Givetian, Middle Devonian) of the Orcadian Basin, Scotland . Scottish Journal of Geology . 57 . sjg2020-009. 10.1144/sjg2020-009 . 224879065 .
Sylvain Adnet . Laurent Marivaux . Henri Cappetta . Anne-Lise Charruault . El Mabrouk Essid . Suzanne Jiquel . Hayet Khayati Ammar . Bernard Marandat . Wissem Marzougui . Gilles Merzeraud . Rim Temani . Monique Vianey-Liaud . Rodolphe Tabuce . 2020 . Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia . Palaeontologia Electronica . 23 . 2 . Article number 23(2):a38 . 10.26879/1085 . 221506440 . free .
Márton Szabó . István Főzy . 2020 . Asteracanthus (Hybodontiformes: Acrodontidae) remains from the Jurassic of Hungary, with the description of a new species and with remarks on the taxonomy and paleobiology of the genus . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 297 . 3 . 295–309 . 10.1127/njgpa/2020/0926 . 225326215 .
Bernard Siguendibo Sambou . Lionel Hautier . Raphael Sarr . Rodolphe Tabuce . Fabrice Lihoreau . Moustapha Thiam . Renaud Lebrun . Jeremy E. Martin . Henri Cappetta . Sylvain Adnet . 2020 . Contribution to the reappraisal of the mid Paleogene ichtyofauna of Western Africa with three new enigmatical elasmobranchs from Thanetian–Lutetian of Senegal . Annales de Paléontologie . 106 . 3 . Article 102400 . 10.1016/j.annpal.2020.102400 . 2020AnPal.10602400S . 216465866 .
Evgeny V. Popov . Marjorie J. Johns . Stephen Suntok . 2020 . A new genus of chimaerid fish (Holocephali, Chimaeridae) from the Upper Oligocene Sooke Formation of British Columbia, Canada . Journal of Vertebrate Paleontology . 40 . 1 . e1772275 . 10.1080/02724634.2020.1772275 . 2020JVPal..40E2275P . 221751740 .
John G. Maisey . Dana J. Ehret . John S.S. Denton . 2020 . A new genus of Late Cretaceous angel shark (Elasmobranchii; Squatinidae), with comments on squatinid phylogeny . American Museum Novitates . 3954 . 1–29 . 10.1206/3954.1 . 2246/7230 . 219700378 .
Giuseppe Marramà . Giorgio Carnevale . Kerin M. Claeson . Gavin J. P. Naylor . Jürgen Kriwet . 2020 . Revision of the Eocene Platyrhina species from the Bolca Lagerstätte (Italy) reveals the first panray (Batomorphii: Zanobatidae) in the fossil record . Journal of Systematic Palaeontology . 18 . 18 . 1519–1542 . 10.1080/14772019.2020.1783380 . 32939187 . 7455076 . 2020JSPal..18.1519M . 221065291 .
Giuseppe Marramà . Giorgio Carnevale . Gavin J. P. Naylor . Massimo Varese . Luca Giusberti . Jürgen Kriwet . 2020 . Anatomy, taxonomy and phylogeny of the Eocene guitarfishes from the Bolca Lagerstätten, Italy, provide new insights into the relationships of the Rhinopristiformes (Elasmobranchii: Batomorphii) . Zoological Journal of the Linnean Society . 192 . 4 . 1090–1110 . 10.1093/zoolinnean/zlaa125 . free .
Linda Frey . Michael I. Coates . Kristen Tietjen . Martin Rücklin . Christian Klug . 2020 . A symmoriiform from the Late Devonian of Morocco demonstrates a derived jaw function in ancient chondrichthyans . Communications Biology . 3 . 1 . Article number 681 . 10.1038/s42003-020-01394-2 . 33203942 . 7672094 .
Alexander O. Ivanov . Merlynd K. Nestell . Galina P. Nestell . Gorden L. Bell Jr. . 2020 . New fish assemblages from the Middle Permian from the Guadalupe Mountains, West Texas, USA . Palaeoworld . 29 . 2 . 239–256 . 10.1016/j.palwor.2018.10.003 . 134496257 .
David J. Cicimurri . Jun A. Ebersole . George Martin . 2020 . Two new species of Mennerotodus Zhelezko, 1994 (Chondrichthyes: Lamniformes: Odontaspididae), from the Paleogene of the southeastern United States . Fossil Record . 23 . 2 . 117–140 . 10.5194/fr-23-117-2020 . 2020FossR..23..117C . 221737251 . free .
Iris Feichtinger . Jürgen Pollerspöck . Mathias Harzhauser . 2020 . A new species of the enigmatic shark genus Nanocetorhinus (Chondrichthyes) from the Oligocene of Austria with palaeoceanographic implications . Austrian Journal of Earth Sciences . 113 . 1–2 . 229–236 . 10.17738/ajes.2020.0014 . 2020AsJES.113..229F . 227313907 . free .
Thomas Reinecke . Jürgen Pollerspöck . Hiroyuki Motomura . Helmut Bracher . Leo Dufraing . Thomas Güthner . Fritz von der Hocht . 2020 . Sawsharks (Pristiophoriformes, Pristiophoridae) in the Oligocene and Neogene of Europe and their relationships with extant species based on teeth and rostral denticles . Palaeontos . 33 .
Oleg A. Lebedev . Alexander O. Ivanov . Valeriy V. Linkevich . 2020 . Chondrichthyan spines from the Famennian (Upper Devonian) of Russia . Acta Geologica Polonica . 70 . 3 . 339–362 . 10.24425/agp.2020.132255 . 219489825 . free .
Plamen S. Andreev . Wenjin Zhao . Nian-Zhong Wang . Moya M. Smith . Qiang Li . Xindong Cui . Min Zhu . Ivan J. Sansom . 2020 . Early Silurian chondrichthyans from the Tarim Basin (Xinjiang, China) . . 15 . 2 . e0228589 . 10.1371/journal.pone.0228589 . 32053606 . 7018067 . 2020PLoSO..1528589A . free .
Samuel L.A. Cooper . David M. Martill . 2020 . A diverse assemblage of pycnodont fishes (Actinopterygii, Pycnodontomorpha) from the mid-Cretaceous, continental Kem Kem beds of South-East Morocco . Cretaceous Research . 112 . Article 104456 . 10.1016/j.cretres.2020.104456 . 216214083 .
Martin Ebert . 2020 . Elongofuro and Altmuehlfuro, new genera of Halecomorphi (Neopterygii) from the Upper Jurassic of the Solnhofen Archipelago and Nusplingen (Germany) . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 295 . 2 . 185–210 . 10.1127/njgpa/2020/0883 . 216400476 .
Yoshitaka Yabumoto . Koji Hirose . Paulo M. Brito . 2020 . A new ichthyodectiform fish, Amakusaichthys goshouraensis gen. et sp. nov. from the Upper Cretaceous (Santonian) Himenoura Group in Goshoura, Amakusa, Kumamoto, Japan . Historical Biology: An International Journal of Paleobiology . 32 . 3 . 362–375 . 10.1080/08912963.2018.1497022 . 2020HBio...32..362Y . 91632554 .
Gary L. Stringer . Werner Schwarzhans . George Phillips . Roger Lambert . 2020 . Highly diversified Late Cretaceous fish assemblage revealed by otoliths (Ripley Formation and Owl Creek Formation, northeast Mississippi, USA) . Rivista Italiana di Paleontologia e Stratigrafia . 126 . 1 . 111–155 . 10.13130/2039-4942/13013 .
Luigi Capasso . 2020 . †Pycnodonts (Neopterygii, †Pycnodontiformes) from the Del Rio Formation (Early Cenomanian, Cretaceous) of Waco Lake, Texas (U.S.A.) . Thalassia Salentina . 42 . 9–24 . 10.1285/i15910725v42p9 .
Alexandre F. Bannikov . Roberto Zorzin . 2020 . A new genus and species of percomorph fish ("stem pleuronectiform") from the Eocene of Bolca in northern Italy . Studi e ricerche sui giacimenti terziari di Bolca, XX - Miscellanea Paleontologica . 17 . 5–14 .
Orangel Aguilera . Ricardo T. Lopes . Felix Rodriguez . Thaís M. dos Santos . Caroline Rodrigues-Almeida . Paulo Almeida . Alessandra S. Machado . Tailan Moretti . 2020 . Fossil sea catfish (Siluriformes; Ariidae) otoliths and in-skull otoliths from the Neogene of the Western Central Atlantic . Journal of South American Earth Sciences . 101 . Article 102619 . 10.1016/j.jsames.2020.102619 . 2020JSAES.10102619A . 219045960 . free .
Jesús Alvarado-Ortega . Bruno Andrés Than-Marchese . María del Pilar Melgarejo-Damián . 2020 . On the Albian occurrence of Armigatus (Teleostei, Clupeomorpha) in America, a new species from the Tlayúa Lagerstätte, Mexico . Palaeontologia Electronica . 23 . 3 . Article number 23(3):a52 . 10.26879/1107 . 229647341 . free .
Melanie Altner . Bettina Reichenbacher . 2020 . A small cichlid species flock from the Upper Miocene (9–10 MYA) of Central Kenya . Hydrobiologia . 848 . 16 . 3613–3637 . 10.1007/s10750-020-04358-z . 221145584 . free .
Robert E. Weems . 2020 . Additions to the bony fish fauna from the early Eocene Nanjemoy Formation of Maryland and Virginia (U.S.A.) . The Mosasaur. The Journal of the Delaware Valley Paleontological Society . XI . 117–152 .
Federico L. Agnolin . Sergio Bogan . 2020 . Goliath catfish Brachyplatystoma Bleeker, 1862 (Siluriformes: Pimelodidae) from the Miocene of Argentina . Journal of South American Earth Sciences . 100 . Article 102551 . 10.1016/j.jsames.2020.102551 . 2020JSAES.10002551A . 216380969 .
Aleksandr Bakaev . Ilja Kogan . 2020 . A new species of Burguklia (Pisces, Actinopterygii) from the Middle Permian of the Volga Region (European Russia) . PalZ . 94 . 1 . 93–106. 10.1007/s12542-019-00487-6 . 2020PalZ...94...93B . 201966465 .
A. F. Bannikov . 2020 . A new genus and species of scombrid fish (Perciformes, Scombroidei, Scombridae) from the Lower Oligocene of the Caucasus . Paleontological Journal . 54 . 1 . 59–67 . 10.1134/S0031030120010037 . 2020PalJ...54...59B . 212936546 .
Kleyton Magno Cantalice Severiano . Jesús Alvarado Ortega . David Bellwood . 2020 . †Chaychanus gonzalezorum gen. et sp. nov.: A damselfish fossil (Pomacentridae; Percomorphaceae), from the Early Paleocene outcrop of Chiapas, Southeastern Mexico . Journal of South American Earth Sciences . 98 . Article 102322 . 10.1016/j.jsames.2019.102322 . 202910177 .
Dmitry Plax . 2020 . A new species of the actinopterygian fish (Osteichthyes, Actinopterygii) from the Eifelian deposits of the Vileyka Buried Ridge (Belarus) . Лiтасфера . 1 . 52 . 68–74 .
Werner Schwarzhans . Rotislav Brzobohatý . Urszula Radwańska . 2020 . Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of the European Gobiidae (Gobiiformes; Teleostei) . Bollettino della Società Paleontologica Italiana . 59 . 2 . 125–173 . 10.4435/BSPI.2020.10 .
Yoshitaka Yabumoto . Mihkail V. Nazarkin . 2020 . Clupea hanishinaensis nomen novum, a replacement name for the Miocene clupeid fish Clupea macrocephala Yabumoto and Nazarkin, 2018 from Nagano, Japan . Paleontological Research . 24 . 3 . 238 . 10.2517/2019PR011 . 220281581 .
Gi-Soo Nam . Mikhail V. Nazarkin . 2020 . A Neogene bristlemouth of the genus Cyclothone (Stomiiformes: Gonostomatidae) from South Korea . Historical Biology: An International Journal of Paleobiology . 33 . 11 . 2639–2645 . 10.1080/08912963.2020.1820000 . 224937137 .
Werner Schwarzhans . Konstantina Agiadi . Giorgio Carnevale . 2020 . Late Miocene–early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance . Rivista Italiana di Paleontologia e Stratigrafia . 126 . 3 . 657–723 . 10.13130/2039-4942/14185 .
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