2020 in paleobotany explained

This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

Flowering plants

Arecales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Echimonocolpites chicxulubensis^[3]	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)			Pollen of a flowering plant, probably a member of the family Arecaceae.
Palmoxylon ceroxyloides^[4]	Sp. nov	In press	Khan, Hazra & Bera in Khan et al.	Late Cretaceous (Maastrichtian)-Paleocene (Danian)	Deccan Intertrappean Beds		A petrified palm stem of a member of the subfamily Ceroxyloideae.
Palmoxylon dindoriensis^[5]	Sp. nov	Valid	Khan, Roy & Bera in Khan et al.	Late Cretaceous (Maastrichtian)-Paleocene (Danian)	Deccan Intertrappean Beds		A petrified palm stem.
Sabalites dawsonii^[6]	Sp. nov	Valid	Greenwood & Conran	Eocene
Sabalites karondiensis^[7]	Sp. nov	In press	Roy, Hazra & Khan in Roy et al.	Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian)	Deccan Intertrappean Beds		A palm frond .
Spinizonocolpites riochiquensis^[8]	Sp. nov	Valid	Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal	Late Cretaceous (Maastrichtian)	Lago Colhué Huapí Formation		A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Cornales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Amersinia littletonensis^[10]	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation
Langtonia parva	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation		A member of the family Mastixiaceae.
Mastixicarpum hoodii	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation		A member of the family Mastixiaceae.
Nyssa gergoei^[11]	Sp. nov	Valid	Hably	Miocene			A tupelo.
Nyssa gyoergyi	Sp. nov	Valid	Hably	Miocene			A tupelo.
Platycrater iljinskajae^[12]	Sp. nov	In press	Denk et al.	Late Oligocene
Portnallia alexanderi	Sp. nov	Valid	Huegele & Manchester	Early Paleocene	Denver Formation		A member of the family Mastixiaceae.

Cucurbitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Coriaripites goodii^[14]	Sp. nov	Valid	Barreda, Palazzesi & Tellería in Renner et al.	Late Cretaceous (Campanian–early Maastrichtian)	Lopez de Bertodano Formation Santa Marta Formation Snow Hill Island Formation	Antarctica	Pollen grains similar to those of extant members of the genus Coriaria.
Echitriporites jolyi	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021.

Ericales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Andrewsiocarpon puryearensis	Sp. nov	Valid	Na, Blanchard & Wang	Middle Eocene	Cockfield Formation		A member of the family Theaceae.
Anubiscarpon^[15]	Gen. et sp. nov	Valid	Smith & Manchester	Middle Eocene	Clarno Formation		A member of the family Theaceae. Genus includes new species A. andersonae.

Fabales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cercis zekuensis^[16]	Sp. nov	Valid	Li et al.	Early Miocene			A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021.
Gleditsia pliocaenica	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene			A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020.
Menendoxylon lutzi	Sp. nov	In press	Baez & Crisafulli	Miocene	Chiquimil		Fossil wood of a member of the family Fabaceae.
Parkiidites marileae^[17]	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021.
Prioria martineziorum	Sp. nov	Valid	Rodríguez-Reyes & Estrada-Ruiz	Oligocene-Miocene	Santiago Formation		A species of Prioria.
Psilastephanocolporites deoliverae	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Psilastephanocolporites endoporatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Announced in 2020; the final version of the article naming it was published in 2021.
Striatopollis grahamii	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)			Pollen of an eudicot, probably a member of the family Fabaceae.

Fagales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Alnus chaybulakensis^[18]	Sp. nov	Valid	Averyanova & Xing	Paleogene			An alder.
Berryophyllum hainanensis^[19]	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		A member of the family Fagaceae.
Carpinus asymmetrica^[20]	Sp. nov	In press	Xue & Jia in Xue et al.	Early Miocene	Maguan		A species of Carpinus.
Carpinus symmetrica	Sp. nov	In press	Xue & Jia in Xue et al.	Early Miocene	Maguan Basin		A species of Carpinus
Carya pipecreekensis^[21]	Sp. nov	In press	Swinehart & Farlow	Late Neogene	Pipe Creek Sinkhole		A hickory.
Castaneophyllum hainanensis	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		A member of the family Fagaceae.
Castaneophyllum lanceolata	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		A member of the family Fagaceae.
Castanopsis bulgarica^[22]	Sp. nov	Valid	Mantzouka, Ivanov & Bozukov	Late Miocene–early Pliocene (late Messinian–early Zanclean)	Pokrovnik		A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021.
Lithocarpus changchangensis	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		A species of Lithocarpus.
Quercus aimeeana^[23]	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus borissovii	Sp. nov	Valid	Averyanova & Xing	Paleogene			An oak.
Quercus campanulata	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus casparyi	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus changchangensis	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		An oak.
Quercus conwentzii	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus longianthera	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus multipilosa	Sp. nov	Valid	Sadowski, Schmidt & Denk	Eocene	Baltic amber	Europe (Baltic Sea region)	An oak.
Quercus paleoargyrotricha	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		An oak.
Quercus paleohypargyrea	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		An oak.
Quercus paleolamellosa	Sp. nov	Valid	Liu & Jin in Liu, Song & Jin	Eocene	Changchang Formation		An oak.

Gentianales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Asclepiadospermum^[24]	Gen. et 2 sp. nov	Valid	Del Rio et al.	Early Eocene	Niubao Formation		An asclepiadoid Apocynaceae genus. Included species A. marginatum and A. ellipticum.
Margocolporites carinae	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021.

Icacinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Iodes elliptica^[25]	Sp. nov	In press	Del Rio et al.	Early Oligocene	Wenshan Basin		A member of the family Icacinaceae.
Iodes passiciensis^[26]	Sp. nov	Valid	Del Rio & De Franceschi	Early Eocene			A member of the family Icacinaceae.
Manchesteria^[27]	Gen. et sp. nov	In press	Stull & Rozefelds in Rozefelds et al.	Cenozoic (mid-Miocene or, more likely, middle Eocene)			A member of the family Icacinaceae. Genus includes new species M. australis.
Pyrenacantha simonsii^[28]	Sp. nov	Valid	Stull et al.	Early Oligocene			A species of Pyrenacantha.

Lamiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Echitricolpites cruziae	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021.
Multiareolites? reticulatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Retistephanocolpites curvimuratus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Announced in 2020; the final version of the article naming it was published in 2021.
Retistephanocolpites pardoi	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021.

Laurales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Actinodaphnoxylon^[29]	Gen. et sp. nov	In press	Akkemik et al.	Eocene (Lutetian)			A member of the family Lauraceae. Genus includes new species A. zileensis.
Mezilaurinoxylon oleiferum^[30]	Sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation		A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Patagonoxylon	Gen. et sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation		A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Thymolepis^[31]	Gen. et sp. nov	Valid	Chambers & Poinar	Late Cretaceous (Cenomanian)	Burmese amber		Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.
Valviloculus^[32]	Gen. et sp. nov	Valid	Poinar et al.	Late Cretaceous (Cenomanian)	Burmese amber		Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis.

Liliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Smilax fujianensis^[33]	Sp. nov	Valid	Dong et al.	Middle Miocene			A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.
Smilax zhangpuensis	Sp. nov	Valid	Dong et al.	Middle Miocene			A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Magnoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Magnolia nanningensis^[34]	Sp. nov	Valid	Huang et al.	Late Oligocene	Nanning Basin		A species of Magnolia.
Magnolia waltheri^[35]	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene			A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020.
Melloniflora^[36]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous	Potomac Group		A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis.

Malvales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Bastardioxylon^[38]	Gen. et sp. nov	In press	Baez & Crisafulli	Miocene	Chiquimil		Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua.
Dipterocarpus dindoriensis^[39]	Sp. nov	Valid	Khan, Spicer & Bera in Khan et al.	Late Cretaceous (Maastrichtian)	Deccan Intertrappean Beds		A species of Dipterocarpus.
Echiperiporites germeraadii	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Announced in 2020; the final version of the article naming it was published in 2021.
Echiperiporites jaramilloi	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021.
Echiperiporites titanicus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021.
Retistephanocolporites elizabeteae	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021.
Veraguasoxylon^[40]	Gen. et sp. nov	Valid	Rodríguez-Reyes & Estrada-Ruiz	Oligocene-Miocene	Santiago Formation		A member of the family Malvaceae. Genus includes new species V. panamense.

Myrtales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Eucalyptus xoshemium^[41]	Sp. nov	Valid	Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon	Eocene (Ypresian)	Huitrera Formation		A species of Eucalyptus.
Mangroveoxylon^[42]	Gen. et comb. nov	In press	Moya & Brea	Late Miocene?	Ituzaingó Formation		A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979).
Myrceugenellites grandiporosum	Sp. nov	Valid	Ruiz, Brea & Pujana in Ruiz et al.	Paleocene (Danian)	Salamanca Formation		A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Primotrapa^[43]	Gen. et sp. nov	Valid	Li & Li in Li et al.	Late Eocene to early Miocene	Hannuoba		A member of Trapoideae. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018).

Oxalidales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cunoniantha^[45]	Gen. et sp. nov	Valid	Jud & Gandolfo	Paleocene (early Danian)	Salamanca Formation		A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021.
Elaeocarpus nanningensis^[46]	Sp. nov	In press	Liu et al.	Late Oligocene	Yongning Formation		A species of Elaeocarpus.
Elaeocarpus prelacunosus	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation		A species of Elaeocarpus.
Elaeocarpus preprunifolioides	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation		A species of Elaeocarpus.
Elaeocarpus prerugosus	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation		A species of Elaeocarpus.
Elaeocarpus preserratus	Sp. nov	In press	Liu et al.	Late Miocene	Foluo Formation		A species of Elaeocarpus.
Elaeocarpus presikkimensis	Sp. nov	In press	Liu et al.	Miocene	Erzitang Formation		A species of Elaeocarpus.

Poales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Monoporopollenites scabratus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Rhizomatites	Gen. et sp. nov	In press	Robledo & Anzótegui in Robledo et al.	Miocene-Pliocene			A member of Cyperaceae. Genus includes new species R. cyperoides.

Proteales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Banksia microphylla^[47]	Sp. nov	Valid	Carpenter in Carpenter & Milne	Late Eocene			A species of Banksia.
Banksieaeidites zanthus	Sp. nov	Valid	Milne in Carpenter & Milne	Late Eocene			A Banksia-like pollen.
Platanus emryi^[48]	Sp. nov	Valid	Huegele, Spielbauer & Manchester	Miocene			A species of Platanus.

Ranunculales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cissampelos defranceschii	Sp. nov	Valid	Del Rio & Su in Del Rio et al.	Middle Eocene	Niubao Formation		A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021.
Clematis csabae	Sp. nov	Valid	Hably	Miocene			A species of Clematis.
Diploclisia praeaffinis^[49]	Sp. nov	Valid	Jia et al.	Late Miocene			A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Menispermites bangorensis^[50]	Sp. nov	Valid	Huang in Del Rio et al.	Middle Eocene	Niubao Formation		A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Menispermites haominae	Sp. nov	Valid	Huang in Del Rio et al.	Middle Eocene	Niubao Formation		A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Menispermites tibetica	Sp. nov	Valid	Huang in Del Rio et al.	Middle Eocene	Niubao Formation		A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Paleoorbicarpum^[51]	Gen. et sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin		A member of the family Menispermaceae. Genus includes new species P. parvum.
Stephania bangorensis	Sp. nov	Valid	Del Rio & Su in Del Rio et al.	Middle Eocene	Niubao Formation		A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.
Stephania geniculata	Sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin		A species of Stephania
Stephania ornamenta	Sp. nov	Valid	Han et al.	Paleocene	Sanshui Basin		A species of Stephania
Stephania shuangxingii	Sp. nov	Valid	Del Rio & Su in Del Rio et al.	Middle Eocene	Niubao Formation		A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Rosales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Berhamniphyllum junrongii^[52]	Sp. nov	Valid	Zhou, Wang & Huang in Zhou et al.	Late Eocene	Markam Basin		A member of the family Rhamnaceae
Crataegus pentagynoides	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene			A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020.
Hemiptelea kryshtofovichii	Sp. nov	Valid	Averyanova & Xing	Paleogene			A member of the family Ulmaceae.
Scabrastephanoporites	Gen. et sp. nov	Valid	Smith et al.	Eocene (Ypresian)			Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

Sapindales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Acer dombeyopsis	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene			A maple. Announced in 2019; the final version of the article naming it was published in 2020.
Acer viburnoides	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene			A maple. Announced in 2019; the final version of the article naming it was published in 2020.
Acer vitiforme	Sp. nov	Valid	Kvaček, Teodoridis & Denk	Pliocene			A maple. Announced in 2019; the final version of the article naming it was published in 2020.
Brosipollis reticulatus	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)			Pollen of a flowering plant, probably a member of the family Burseraceae.
Choerospondias fujianensis^[53]	Sp. nov	In press	Wang et al.	Miocene			A species of Choerospondias.
Llanodelacruzoxylon^[54]	Gen. et sp. nov		Rodríguez-Reyes, Estrada-Ruiz & Gasson	Oligocene–Miocene	Santiago Formation		A member of the family Anacardiaceae. Genus includes new species L. sandovalii.
Manchestercarpa^[55]	Gen. et sp. nov	Valid	Atkinson	Late Cretaceous (Campanian)			A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.
Parametopioxylon^[56]	Gen. et sp. nov	Valid	Franco et al.	Miocene	Ituzaingó Formation		A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.
Psilastephanocolporites hammenii	Sp. nov	Valid	Smith et al.	Eocene (Ypresian)			Pollen of a flowering plant, probably a member of the family Meliaceae
Quinquala^[57]	Gen. et sp. nov	Valid	Manchester & Disney in Manchester, Disney & Pham	Eocene	Clarno Formation Tepee Trail Formation	( )	A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata.
Rousea cavitata	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Announced in 2020; the final version of the article naming it was published in 2021.

Saxifragales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Corylopsis grisea^[58]	Sp. nov	Valid	Quirk & Hermsen	Early Pliocene	Gray Fossil Site		A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021.
Itea polyneura^[59]	Sp. nov	In press	Huang & Tian in Tian et al.	Oligocene	Huazhige Formation		A species of Itea.
Protoaltingia^[60]	Gen. et sp. nov	Valid	Scharfstein, Stockey & Rothwell	Late Cretaceous (Coniacian)			A member of the family Altingiaceae. Genus includes new species P. comoxense.

Trochodendrales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Eotrochion^[62]	Gen. et sp. nov	Valid	Manchester, Kvaček & Judd	Paleocene			A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021.
Paraconcavistylon	Gen. et comb. nov	Valid	Manchester, Kvaček & Judd	Ypresian			A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021.
Trochodendron infernense	Sp. nov	Valid	Manchester, Kvaček & Judd	Paleocene			A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021.

Other angiosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aextoxicoxylon kawasianus^[64]	Sp. nov	Valid	Vera et al.	Late Cretaceous	Puntudo Chico Formation		A fossil dicot wood
Atlantocarpus^[65]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Albian)	Potomac Group		An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis.
Carpolithes gergoei	Sp. nov	Valid	Hably & Erdei in Hably	Miocene			A fossil fruit of a flowering plant of uncertain phylogenetic placement.
Catanthus^[66]	Gen. et sp. nov	In press	Friis, Crane & Pedersen	Early Cretaceous			An early flowering plant. Genus includes new species C. dolichostemon.
Cavilignum^[67]	Gen. et sp. nov	Valid	Siegert & Hermsen	Early Pliocene	Gray Fossil Site		A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii.
Chainandra^[68]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber		Genus includes new species C. zeugostylus.
Cichoreacidites? flammulatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Crotonoidaepollenites echinatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Cyathitepala^[69]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber		Genus includes new species C. papillosa.
Dasykothon^[70]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber		A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.
Dinganthus^[71]	Gen. et sp. nov	Valid	Liu et al.	Miocene	Dominican amber		A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera.
Echistephanoporites annulatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Eofructus^[72]	Gen. et sp. nov	Valid	Han & Wang	Early Cretaceous	Yixian Formation		An infructescence including a central axis and five fruits resembling Liaoningfructus. Genus includes new species E. liutiaogouensis.
Foveotricolporites crassus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Inaperturopollenites microechinatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Ladakhipollenites carmoi	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Lambertiflora	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Albian)	Potomac Group		An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans.
Malvacipolloides diversus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Malvacipolloides echibaculatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Malvacipolloides romeroae	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Menatanthus^[73]	Gen. et sp. nov	Valid	Uhl, Paudayal & El Atfy in Uhl et al.	Paleocene (Thanetian)			A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021.
Mugideiriflora	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Aptian-early Albian)	Almargem Formation		An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica.
Phantophlebia^[74]	Gen. et sp. nov	Valid	Poinar & Chambers	Late Cretaceous (Cenomanian)	Burmese amber		A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.
Platanites willigeri^[75]	Sp. nov	Valid	Halamski & Kvaček in Halamski et al.	Late Cretaceous (Santonian)	Czerna Formation		Trifoliolate platanoid leaves.
Psilaperiporites delicatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Psilaperiporites lunaris	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Ranunculacidites reticulatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Rasenganus^[76]	Gen. et sp. nov	Valid	Xing & Gu	Late Cretaceous (Cenomanian)	Burmese amber		A possible epizoochorous fruit. Genus includes new species R. auricularus.
Retibrevitricolpites microreticulatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retibrevitricolporites costaporus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retibrevitricolporites? toigoae	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retipollenites solimoensis	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Retitriporites crassoreticulatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Rhoipites alfredii	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.
Singpuria^[77]	Gen. et sp. nov	Valid	Ramteke, Manchester & Nagrale in Ramteke et al.	Late Cretaceous (Maastrichtian)	Deccan Intertrappean Beds		A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei.
Sinoherba^[78]	Gen. et sp. nov	Valid	Liu & Wang in Liu, Chen & Wang	Early Cretaceous (Barremian–Aptian)	Yixian Formation		An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021.
Varifructus^[79]	Gen. et sp. nov	In press	Liu et al.	Early Cretaceous	Yixian Formation		An early flowering plant. Genus includes new species V. lingyuanensis.
Wireroadia^[80]	Gen. et sp. et comb. nov	Valid	Zhang et al.	Late Cretaceous (Cenomanian to Santonian)		( )	A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick).

Pinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Agathis ledongensis^[81]	Sp. nov	Valid	Oskolski et al.	Late Oligocene–early Miocene	Qiutangling Formation		A species of Agathis
Agathoxylon cozzoi^[82]	Sp. nov	Valid	Gnaedinger & Zavattieri	Late Triassic	Chihuido Formation
Araucaria cuneoi^[83]	Sp. nov	Valid	Noll & Kunzmann	Middle Jurassic			A species of Araucaria.
Araucaria famii^[84]	Sp. nov	Valid	Stockey & Rothwell	Late Cretaceous (Campanian)			A species of Araucaria.
Araucaria fildesensis^[85]	Sp. nov	Valid	Shi et al.	Eocene	Fossil Hill Formation	Antarctica (King George Island)	A species of Araucaria
Araucaria huncoensis^[86]	Sp. nov	Valid	Rossetto-Harris in Rossetto-Harris et al.	Early Eocene	Laguna del Hunco Formation		A species of Araucaria.
Araucaria stockeyana	Sp. nov	Valid	Noll & Kunzmann	Middle Jurassic			A species of Araucaria.
Araucarites pachacuteci^[87]	Sp. nov	In press	Martínez in Martínez et al.	Early Cretaceous (Berriasian–Valanginian)	Huancané Formation		A member of the family Araucariaceae
Brachyoxylon zhouii^[88]	Sp. nov	Valid	Jiang et al.	Early Cretaceous	Guantou Formation		A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021.
Brachyphyllum sattlerae^[89]	Sp. nov	Valid	Batista et al.	Early Cretaceous (Aptian)	Crato Formation		A member of the family Pinidae.
Callialastrobus^[90]	Gen. et sp. nov	In press	Kvaček & Mendes	Early Cretaceous (late Aptian–early Albian)	Lusitanian Basin		A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai.
Cedrus anatolica^[91]	Sp. nov	Valid	Akkemik	Early Miocene	Hançili Formation		A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021.
Circoporoxylon tibetense^[92]	Sp. nov	In press	Xia et al.	Middle Jurassic (Callovian)	Xiali Formation		Possibly a member of the family Podocarpaceae.
Comoxostrobus^[93]	Gen. et sp. nov	Valid	Stockey, Rothwell & Atkinson	Late Cretaceous (early Coniacian)			A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii.
Cupressinoxylon klimovii^[94]	Nom. nov	Valid	Blokhina	Miocene			A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989).
Cupressinoxylon llantenesense	Sp. nov	Valid	Gnaedinger & Zavattieri	Late Triassic	Llantenes Formation
Cupressinoxylon manuelii^[95]	Sp. nov	In press	Ríos-Santos, Cevallos-Ferriz & Pujana	Late Cretaceous (Campanian-Maastrichtian)	Cabullona Group
Ductoagathoxylon wangii^[96]	Sp. nov	In press	Gou & Feng in Gou et al.	Middle Jurassic	Xishanyao Formation		A conifer stem.
Friisia^[97]	Gen. et sp. nov	In press	Mendes & Kvaček	Early Cretaceous (late Aptian – early Albian)	Lusitanian Basin		An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.
Juniperoxylon acarcaea^[98]	Sp. nov	In press	Akkemik	Early Miocene	Hançili Formation		A member of the family Cupressaceae.
Lesbosoxylon kemaliyensis^[99]	Sp. nov	Valid	Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım	Miocene	Divriği Formation		A member of the family Pinaceae.
Marskea cuspidata^[100]	Sp. nov	Valid	Frolov & Mashchuk	Middle Jurassic	Prisayan Formation		A member of the family Taxaceae.
Mukawastrobus^[101]	Gen. et sp. nov	Valid	Stockey, Nishida & Rothwell	Late Cretaceous (late Campanian—early Maastrichtian)			A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi.
Piceoxylon yumeniense^[102]	Sp. nov	Valid	Zhou, Peng, Deng, Zhang & Yang in Zhou et al.	Early Cretaceous	Xiagou Formation		Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021.
Pinuxylon selmeierianum^[103]	Sp. nov	Valid	Dolezych & Reinhardt	Paleogene	Eureka Sound Group		A member of the family Pinaceae described on the basis of fossil wood
Protophyllocladoxylon chijinense	Sp. nov	Valid	Zhou, Peng, Deng, Zhang & Yang in Zhou et al.	Early Cretaceous	Xiagou Formation		Announced in 2020; the final version of the article naming it was published in 2021.
Tsuga asiatica^[104]	Sp. nov	Valid	Wu & Zhou in Wu et al.	Late Paleogene			A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020.

Other seed plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Amyelon turpanense^[105]	Sp. nov	In press	Shi, Yu & Sun	Permian (Lopingian)	Wutonggou		A root of a member of Cordaitales
Androstrobus obovatus^[106]	Sp. nov	In press	Bodnar et al.	Late Triassic	Potrerillos Formation		A member of Cycadales.
Araripestrobus^[107]	Gen. et sp. nov	Valid	Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al.	Early Cretaceous (Aptian-Albian)	Crato		A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus.
Archaeopetalanthus^[108]	Gen. et sp. nov	Valid	Naugolnykh	Carboniferous	Listvjanskaya		A member of Pinophyta belonging to the group Vojnovskyales. Genus includes new species A. progressus.
Battenispermum^[109]	Gen. et sp. nov	In press	Mendes, Pedersen & Friis	Early Cretaceous			A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum.
Carpolithus volantus^[110]	Sp. nov	Valid	Gómez et al.	Early Cretaceous (Aptian)	La Cantera Formation		A fossil seed, possibly produced by a member of Gnetales.
Ductolobatopitys^[111]	Gen. et sp. nov	Valid	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation		A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.
Filigigantopteris^[112]	Gen. et 2 sp. nov	In press	Zhou et al.	Late Permian	Nayixiong Formation		A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia.
Ginkgo pediculata^[113]	Sp. nov	In press	Deng, Yang & Zhou	Early Cretaceous			A species of Ginkgo.
Jianchangia^[114]	Gen. et sp. nov	Valid	Yang, Wang & Ferguson	Early Cretaceous (Aptian)	Jiufotang Formation		A member of Ephedraceae. Genus includes new species J. verticillata.
Johniphyllum^[115]	Gen. et sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)			A member of Voltziales. Genus includes new species J. multinerve.
Jordaniopteris^[116]	Gen. et comb. nov	Valid	Anderson in Anderson et al.	Permian (possibly Lopingian)	Um Irna Formation		A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).
Kaokoxylon brasiliensis	Sp. nov	Valid	Conceição, Neregato & Iannuzzi in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation		A conifer described on the basis of fossil wood.
Nilssonia mirovanae^[117]	Sp. nov	Valid	Čepičková & Kvaček	Late Cretaceous (Cenomanian)	Peruc-Korycany Formation		A cycad.
Novaiorquepitys^[118]	Gen. et sp. nov	In press	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation		A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.
Ovalocarpus butmanii^[119]	Sp. nov	Valid	Naugolnykh & Linkevich	Permian (Artinskian)			A member of Ginkgoales belonging to the family Cheirocladaceae.
Palaeocupressinoxylon^[120]	Gen. et sp. nov	In press	Wan, Yang & Wang	Late Permian	Turpan–Hami Basin		A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.
Phoenicopsis anadyrensis^[121]	Sp. nov	Valid	Nosova in Zolina et al.	Late Cretaceous–Paleocene (Maastrichtian–Danian)	Rarytkin		A member of Czekanowskiales
Pseudovoltzia sapflorensis	Sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)			A member of Voltziales
Pterostoma neehoffii^[122]	Sp. nov	Valid	Conran et al.	Middle Miocene			A cycad
Pteruchus frenguellii^[123]	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation		A pollen organ of a seed fern
Pteruchus lepidus	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation		A pollen organ of a seed fern
Samaropsis jinchangensis^[124]	Sp. nov	Valid	Hua & Sun in Hua et al.	Early Permian			A seed fossil. Announced in 2019; the final version of the article naming was published in 2020.
Umaltolepis involuta^[125]	Sp. nov	In press	Nosova	Middle Jurassic	Angren Formation
Umaltolepis sogdianica	Sp. nov	In press	Nosova	Middle Jurassic	Angren Formation
Umkomasia aequatorialis	Sp. nov	In press	Blomenkemper et al.	Late Permian	Umm Irna Formation		A cupulate structure of a seed fern
Wantus	Gen. et sp. nov	Valid	Looy & Duijnstee	Permian (Guadalupian)			A member of Voltziales. Genus includes new species W. acaulis.
Wudaeophyton^[126]	Gen. et sp. nov	Valid	Pšenička et al.	Early Permian	Taiyuan Formation		A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.
Yangopteris^[127]	Gen. et comb. nov	In press	Zhou et al.	Permian (Asselian)			A seed fern; a new genus for "Alethopteris" ascendens Halle.
Yvyrapitys	Gen. et sp. nov	In press	Conceição & Crisafulli in Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation		A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

Other plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Annularia paisii^[128]	Sp. nov	Valid	Correia et al.	Carboniferous (Gzhelian)
Blasiites huolinhensis^[129]	Sp. nov	In press	Li et al.	Early Cretaceous	Huolinhe Formation		A liverwort belonging to the family Blasiaceae.
Botryopteris multifolia^[130]	Sp. nov	In press	He et al.	Permian (Lopingian)	Junlian Formation		A fern
Birisia mandshurica^[131]	Sp. nov	Valid	Golovneva, Grabovskiy & Zolina	Early Cretaceous (Albian)	Frentsevka Formation		A fern belonging to the family Dicksoniaceae.
Calamites cambrensis^[132]	Sp. nov	Valid	Thomas	Carboniferous (Pennsylvanian)
Catenuporella^[133]	Gen. et sp. nov	Valid	Zhang et al.	Late Ordovician	Ordos Basin		A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia.
Charaxis spicatus^[134]	emend. nov.	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Maestrat Basin		Thallus of Echinochara lazarii. Both taxa has been firstly found anatomically attached
Chlamydomonas hanublikanus^[135]	Sp. nov		Vršanská & Hinkelman	Late Cretaceous (Cenomanian)	Burmese amber		A species of Chlamydomonas
Circinites^[136]	Gen. et sp. nov	In press	Robledo & Anzótegui in Robledo et al.	Miocene-Pliocene			A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.
Clavatisporites cenomaniana	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla Formation		Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Clavator calcitrapus var. jiangluoensis	comb. nov	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Maestrat Basin		Clavatoracean species.
Collarecodium? nezpercae	Sp. nov	Valid	Bucur & Rigaud in Bucur et al.	Late Triassic (Norian)			A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.
Collarisporites minor	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla Formation		Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
Coniopteris sandaolingensis^[137]	Sp. nov	Valid	Yuan & Sun in Yuan et al.	Middle Jurassic	Xishanyao Formation
Dimicheleodendron^[138]	Gen. et comb. nov	Valid	Thomas & Cleal	Carboniferous			A lycophyte; a new genus for "Lepidodendron" hickii.
Drynaria diplosticha^[139]	Sp. nov	Valid	Yu & Xie in Yu et al.	Late Miocene	Bangmai Formation		A fern belonging to the family Polypodiaceae.
Echinochara lazarii	comb. nov	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Maestrat Basin		A member of Clavatoraceans.
Equicalastrobus pusillus^[140]	Sp. nov	Valid	Zhang & Yan in Zhang et al.	Late Triassic	Baojishan Basin		A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021.
Equisetum yenbaiense^[141]	Sp. nov	Valid	Aung et al.	Late Miocene			A species of Equisetum
Equisetum yongpingense	Sp. nov	Valid	Aung et al.	Late Pliocene	Sanying Formation		A species of Equisetum
Filippoporella^[142]	Gen. et sp. nov	In press	Sokač & Grgasović	Early Paleocene			A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi.
Frullania partita^[143]	Sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber		A liverwort, a species of Frullania
Frullania vanae^[144]	Sp. nov	Valid	Mamontov et al.	Eocene	Rovno amber		A liverwort, a species of Frullania
Gippslandites^[145]	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Late Silurian–early Devonian			A member of Zosterophyllaceae. Genus includes new species G. minutus.
Gmujij^[146]	Gen. et sp. nov	Valid	Pfeiler & Tomescu	Devonian (Emsian)	Battery Point		An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021.
Griphoporella minuta	Sp. nov	Valid	Bucur & Peybernes in Bucur et al.	Late Triassic			A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Hansopteris^[147]	Gen. et sp. nov	In press	Zhou et al.	Permian (Asselian)			An anachoropterid fern. Genus includes new species H. uncinatus.
Holosporella magna	Sp. nov	Valid	Bucur & Fucelli in Bucur et al.	Late Triassic (Norian)			A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella? rossanae^[148]	Sp. nov	Valid	Bucur & Del Piero in Bucur et al.	Late Triassic (Norian)			A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Inocladus^[149]	Gen. et comb. nov	Valid	LoDuca et al.	Silurian			An alga related to the group Bryopsidales. Genus includes "Buthotrephis" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021.
Intermurella ordosensis	Sp. nov	Valid	Zhang et al.	Late Ordovician	Ordos Basin		A green alga belonging to the group Dasycladales.
Jurafructus^[150]	Gen. et sp. nov	Valid	Chen et al.	Middle−Late Jurassic	Jiulongshan Formation		A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.
Keraphyton^[151]	Gen. et sp. nov	Valid	Champreux, Meyer-Berthaud & Decombeix	Devonian (Famennian)	Mandowa Mudstone Formation		A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.
Khasurtya^[152]	Gen. et sp. nov	Valid	Mamontov in Kopylov et al.	Early Cretaceous			A liverwort belonging to the group Marchantiidae. Genus includes new species K. ginkgoides.
Lepidodendron demkinae^[153]	Sp. nov	Valid	Mosseichik	Carboniferous (Viséan)
Lobatannularia linjiaensis^[154]	Sp. nov	Valid	Xu et al.	Middle Triassic	Linjia Formation		A member of Equisetales.
Lygodium sanshuiense^[155]	Sp. nov	Valid	Naugolnykh et al.	Paleocene	Buxin Formation		A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020.
Metzgeriites kujiensis^[156]	Sp. nov	Valid	Katagiri in Katagiri & Shinden	Late Cretaceous (Santonian)	Tamagawa Formation		A liverwort.
Munieria martinclosasi	comb. nov.	Valid	Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.	Barremian	Lebanon		Clavatoracean thallus. Formerly known as Charaxis martinclosasi
Neoarthropitys^[157]	Gen. et sp. nov	In press	Gnaedinger et al.	Middle Triassic	Quebrada de los Fósiles		A member of Equisetales. Genus includes new species N. gondwanaensis.
Osmundacaulis asiatica^[158]	Sp. nov	Valid	Cheng et al.	Cretaceous			A member of the family Osmundaceae
Osmundacaulis sinica	Sp. nov	Valid	Cheng et al.	Cretaceous			A member of the family Osmundaceae
Ovoidites circumplicatus^[159]	Sp. nov	Valid	Zavattieri, Gutiérrez & Monti	Middle Triassic	Quebrada de los Fósiles Formation		A green alga belonging to the group Zygnematales.
Ovoidites tripartitus	Sp. nov	Valid	Zavattieri, Gutiérrez & Monti	Middle Triassic	Quebrada de los Fósiles Formation		A green alga belonging to the group Zygnematales.
Palaeostachya guanglongii^[160]	Sp. nov	In press	Liu et al.	Permian (Asselian)	Taiyuan Formation		A member of the family Calamitaceae.
Parazosterophyllum	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Late Silurian–early Devonian			A member of Zosterophyllaceae. Genus includes new species P. timsiae.
Patruliuspora oregonica	Sp. nov	Valid	Bucur & Rigaud in Bucur et al.	Late Triassic (Norian)			A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Patruliuspora pacifica	Sp. nov	Valid	Bucur, Del Piero & Peyrotty in Bucur et al.	Late Triassic (Norian)			A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Pellites hamiensis^[161]	Sp. nov	Valid	Li et al.	Middle Jurassic	Xishanyao		A liverwort belonging to the family Pelliaceae.
Plenasium (Aurealcaulis) elegans^[162]	Sp. nov	In press	Hiller et al.	Eocene	Na Duong Formation		A member of Osmundaceae
Pleuromeia shaolinii^[163]	Sp. nov	Valid	Zhang & Wang in Zhang et al.	Middle Triassic	Linjia
Polycingulatisporites multiverrucata^[164]	Sp. nov	Valid	Santamarina in Santamarina et al.	Late Cretaceous (Cenomanian)	Mata Amarilla		Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020.
Polypodiisporites minutiverrucatus	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.
Polysporia baetica^[165]	Sp. nov	In press	Álvarez-Vázquez, Bek & Drábková	Carboniferous (Pennsylvanian)	Peñarroya-Belmez-Espiel Coalfield		A member of Isoetales
Polystichum pacltovae^[166]	Sp. nov	Valid	Kvaček in Kvaček & Teodoridis	Oligocene			A fern, a species of Polystichum
Proodontosoria^[167]	Gen. et sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber		A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.
Proterocladus antiquus^[168]	Sp. nov	Valid	Tang et al.	Mesoproterozoic circa 1 Ga	Nanfen		An early siphonocladalean chlorophyte
Psilatriletes cozzuolii	Sp. nov	Valid	Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito	Miocene	Solimões Basin		Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.
Psilochara monevaensis^[169]	Sp. nov	Valid	Sanjuan & Soulié-Märsche	Middle Miocene			A charophyte.
Qianshouia^[170]	Gen. et sp. nov	Valid	Huang et al.	Late Devonian	Wutong Formation		A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira.
Scolecopteris minuta^[171]	Sp. nov	In press	Wan et al.	Early Permian	Taiyuan Formation		A fern belonging to the group Marattiales.
Sigillaria pfefferkornii^[172]	Sp. nov	In press	D'Antonio, Boyce & Wang	Permian (Asselian)
Sigillaria wudensis	Sp. nov	In press	D'Antonio, Boyce & Wang	Permian (Asselian)
Sphaerochara miocenica	Sp. nov	Valid	Sanjuan & Soulié-Märsche	Miocene			A charophyte.
Thyrsopteris cretacea^[173]	Sp. nov	Valid	Li et al.	Late Cretaceous (Cenomanian)	Burmese amber		A species of Thyrsopteris
Tomiostrobus sinensis^[174]	Sp. nov	Valid	Feng in Feng et al.	Early Triassic (Induan)	Kayitou Formation		A member of the family Isoetaceae.
Tumidopteris astra^[175]	Sp. nov	Valid	Naugolnykh	Permian (Roadian)	Pechora coal basin		A fern belonging to the family Gleicheniaceae.
Ufadendron elongatum^[176]	Sp. nov	Valid	Tang et al.	Late Permian	Linxi		A lycopsid belonging to the family Tomiodendraceae
Uzhurodendron^[177]	Gen. et sp. nov	Valid	Mosseichik & Filimonov	Carboniferous (Tournaisian)	Bystrianskaya		A member of Lycopodiopsida. Genus includes new species U. asiaticum.
Zeilleria fosteri^[178]	Sp. nov	Valid	Thomas et al.	Carboniferous (Bashkirian)			A fern

General research

• A study on the evolutionary history of green plants is published by Nie et al. (2020).^[179]
• Description of new fossil material of Yurtusia uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.^[180]
• A study on the phylogenetic relationships of extant and fossil complex thalloid liverworts (Marchantiidae) is published by Flores et al. (2020).^[181]
• Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.^[182]
• An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).^[183]
• A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).^[184]
• A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.^[185]
• A study on the architecture and development of the Carboniferous arborescent lycopsid Paralycopodites is published by DiMichele & Bateman (2020).^[186]
• New information on the anatomy of Weichselia reticulata is presented by Blanco-Moreno, Decombeix & Prestianni (2020).^[187]
• A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).^[188]
• New information on the morphology of Paleoazolla patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.^[189]
• A study aiming to determine which ferns were most likely to be the producers of Cyathidites spores from earliest Paleocene plant localities across western North America, and were most likely to be among the first plants in western North America to thrive in the immediate aftermath of the Cretaceous–Paleogene extinction event, is published by Berry (2020).^[190]
• A study on the morphology and development of Genomosperma, and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).^[191]
• A pollen organ resembling seed fern pollen organs Dictyothalamus and Melissiotheca is described from the Lopingian Umm Irna Formation (Jordan) by Zavialova et al. (2020), who interpret this finding as evidence of persistence of lyginopterid seed ferns until the late Permian.^[192]
• Evidence of increasing atmospheric CO₂ concentration at the onset of the end-Triassic extinction event, based on data from fossil leaves of the seed fern Lepidopteris ottonis from southern Sweden, is presented by Slodownik, Vajda & Steinthorsdottir (2020), who confirm L. ottonis as a valid proxy for pCO₂ reconstructions.^[193]
• A study on the anatomy of the seed cone scales of Krassilovia mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.^[194]
• A study on the microscopic wood anatomy of a fossil tree trunk of Agathoxylon arizonicum with the characteristic external features of a fire scar from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) is published by Byers et al. (2020), who evaluate the implications of this specimen for the knowledge of the evolution of fire-adapted plant traits.^[195]
• A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.^[196]
• A study on evolutionary history of conifers as indicated by fossil and molecular data, aiming to determine whether the rise of angiosperms drove the decline of conifers and other gymnosperms, is published by Condamine et al. (2020).^[197]
• Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).^[198]
• Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.^[199]
• A study on the morphology of palm and palm-like pollen from the Eocene Yaw Formation (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).^[200]
• Fossils fruits of Illigera eocenica, representing the second fossil occurrence of Illigera worldwide and the first in Asia, are described from the Eocene Niubao Formation (central Tibetan Plateau) by Wang et al. (2020), who evaluate the implications of this finding for the knowledge of the climate in the central Tibetan Plateau during the early middle Eocene, and for the knowledge of the floristic links between Asia and North America during the Paleogene.^[201]
• A study on the morphology and phylogenetic relationships of Montsechia vidalii is published by Gomez et al. (2020).^[202]
• Eocene leaves of members of the family Urticaceae with stinging trichomes are described from the Okanogan Highlands (British Columbia, Canada) by DeVore et al. (2020).^[203]
• A revision of the fossil record of the family Nothofagaceae from South America is published by Pujana et al. (2020).^[204]
• A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).^[205]
• A study on the impact of ecological disturbances around the Permian–Triassic boundary (from the Wuchiapingian to Ladinian) on land plant communities is published by Nowak, Vérard & Kustatscher (2020).^[206]
• A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment is published by Mathewes, Greenwood & Love (2020).^[207]
• Evidence from Eocene plant fossils from the Bangong-Nujiang suture indicating that the Tibetan Plateau area hosted a diverse subtropical ecosystem approximately 47 million years ago and that this area was both low and humid at the time is presented by Su et al. (2020), who also report that the composition of this flora is similar to Early-Middle Eocene floras in both North America and Europe, but shows little affinity to Eocene floras from the Indian Plate.^[208]
• A study aiming to estimate leaf dry mass per area in fossil plants from 22 western North American sites spanning the Eocene–Oligocene transition is published online by Butrim & Royer (2020), who evaluate the implications of their findings for the knowledge of the impact of the environmental changes occurring during the Eocene–Oligocene transition on leaf-economic strategies of plants.^[209]
• A study on the Neogene paleobotanical record and climate in the northernmost part of the Central Andean Plateau, based on data from the Descanso Formation (Peru), is published by Martínez et al. (2020), who report the earliest evidence of a puna-like ecosystem in the Pliocene and a montane ecosystem without modern analogs in the Miocene.^[210]
• Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (≡ Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020.^[211] This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
• The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020,^[212] with implications for the evolution of leaf shape in the early radiation of the angiosperms.
• A study on the phylogenetic relationships of 10 Cretaceous flower taxa (Chloranthistemon endressii, Dakotanthus cordiformis, Kajanthus lusitanicus, Mauldinia mirabilis, Microvictoria svitkoana, Paleoclusia chevalieri, Paradinandra suecica, Spanomera mauldiniensis, Tylerianthus crossmanensis and Virginianthus calycanthoides) is published by Schönenberger et al. (2020).^[213]

Notes and References

1. Shook Ling Low . Tao Su . Teresa E. V. Spicer . Fei-Xiang Wu . Tao Deng . Yao-Wu Xing . Zhe-Kun Zhou . 2020 . Oligocene Limnobiophyllum (Araceae) from the central Tibetan Plateau and its evolutionary and palaeoenvironmental implications . Journal of Systematic Palaeontology . 18 . 5 . 415–431 . 10.1080/14772019.2019.1611673 . 2020JSPal..18..415L . 208589882 .
2. Yuling Na . Jane Blanchard . Hongshan Wang . 2020 . Fruits, seeds and flowers from the Puryear clay pit (middle Eocene Cockfield Formation), western Tennessee, USA . Palaeontologia Electronica . 23 . 3 . Article number 23(3):a49 . 10.26879/1045 . free .
3. Vann Smith . Sophie Warny . David M. Jarzen . Thomas Demchuk . Vivi Vajda . Sean P.S. Gulick . 2020 . Paleocene–Eocene palynomorphs from the Chicxulub impact crater, Mexico. Part 2: angiosperm pollen . Palynology . 44 . 3 . 489–519 . 10.1080/01916122.2019.1705417 . 2020Paly...44..489S . 213827225 .
4. Mahasin Ali Khan . Manoshi Hazra . Sumana Mahato . Robert A. Spicer . Kaustav Roy . Taposhi Hazra . Manosij Bandopadhaya . Teresa E.V. Spicer . Subir Bera . 2020 . A Cretaceous Gondwana origin of the wax palm subfamily (Ceroxyloideae: Arecaceae) and its paleobiogeographic context . Review of Palaeobotany and Palynology . 283 . Article 104318 . 10.1016/j.revpalbo.2020.104318 . 2020RPaPa.28304318K . 224946279 .
5. Mahasin Ali Khan . Kaustav Roy . Taposhi Hazra . Sumana Mahato . Subir Bera . 2020 . A new coryphoid palm from the Maastrichtian-Danian sediments of Madhya Pradesh and its palaeoenvironmental implications . Journal of the Geological Society of India . 95 . 1 . 75–83 . 10.1007/s12594-020-1388-1 . 2020JGSI...95...75K . 210134584 .
6. David Robert Greenwood . John G. Conran . 2020 . Fossil coryphoid palms from the Eocene of Vancouver, British Columbia, Canada . International Journal of Plant Sciences . 181 . 2 . 224–240 . 10.1086/706450 . 208587364 .
7. Kaustav Roy . Taposhi Hazra . Manoshi Hazra . Sumana Mahato . Subir Bera . Mahasin Ali Khan . 2020 . A new coryphoid costapalmate palm leaf from the Maastrichtian-Danian of India . Botany Letters . 168 . 2 . 155–166 . 10.1080/23818107.2020.1845974 . 229408918 .
8. Patricia Vallati . Andrea De Sosa Tomas . Gabriel Casal . 2020 . A Maastrichtian terrestrial palaeoenvironment close to the K/Pg boundary in the Golfo San Jorge basin, Patagonia, Argentina . Journal of South American Earth Sciences . 97 . Article 102401 . 10.1016/j.jsames.2019.102401 . 2020JSAES..9702401V . free .
9. M. Laura Pipo . Ari Iglesias . Josefina Bodnar . 2020 . A new vesselless angiosperm stem with a cambial variant from the Upper Cretaceous of Antarctica . Acta Palaeontologica Polonica . 65 . 2 . 261–272 . 10.4202/app.00697.2019 . free . 11336/136234 . free .
10. Indah B. Huegele . Steven R. Manchester . 2020 . An early Paleocene carpoflora from the Denver Basin of Colorado, USA, and its implications for plant-animal interactions and fruit size evolution . International Journal of Plant Sciences . 181 . 6 . 646–665 . 10.1086/707727 . 219748982 . free .
11. Lilla Hably . 2020 . The Karpatian (late early Miocene) flora of the Mecsek area . Acta Palaeobotanica . 60 . 1 . 51–122 . 10.35535/acpa-2020-0003 . free .
12. Thomas Denk . Johannes Martin Bouchal . Pavel Smirnov . Yaroslav Trubin . 2020 . Late Oligocene leaf and pollen flora of Southwestern Siberia: taxonomy, biogeography and palaeoenvironments . Historical Biology: An International Journal of Paleobiology . 33 . 11 . 2951–2976 . 10.1080/08912963.2020.1839064 . 230547821 .
13. Hai Zhu . Steven R. Manchester . 2020 . Fruit of Staphylea (Staphyleaceae) from the Oligocene of Montana, USA . Review of Palaeobotany and Palynology . 280 . Article 104275 . 10.1016/j.revpalbo.2020.104275 . 2020RPaPa.28004275Z . 225729365 . free .
14. Susanne S. Renner . Viviana D. Barreda . María Cristina Tellería . Luis Palazzesi . Tanja M. Schuster . 2020 . Early evolution of Coriariaceae (Cucurbitales) in light of a new early Campanian (ca. 82 Mya) pollen record from Antarctica . Taxon . 69 . 1 . 87–99 . 10.1002/tax.12203 . free .
15. MacKenzie Allan Smith . Steven R. Manchester . 2020 . CT-scans of capsules from the Clarno Formation (Oregon, USA) reveal an extinct Eocene theaceous taxon . Acta Palaeobotanica . 60 . 2 . 251–258 . 10.35535/acpa-2020-0013 . free .
16. Xiang-Chuan Li . Steven R. Manchester . Qin Wang . Liang Xiao . Tian-Long Qi . Yun-Zhi Yao . Dong Ren . Qiang Yang . 2020 . A unique record of Cercis from the late early Miocene of interior Asia and its significance for paleoenvironment and paleophytogeography . Journal of Systematics and Evolution . 59 . 6 . 1321–1338 . 10.1111/jse.12640 . 219523602 . free .
17. Fátima Praxedes Rabelo Leite . Silane Aparecida Ferreira da Silva-Caminha . Carlos D'Apolito . 2020 . New Neogene index pollen and spore taxa from the Solimões Basin (Western Amazonia), Brazil . Palynology . 45 . 1 . 115–141 . 10.1080/01916122.2020.1758971 . 219090032 .
18. A.L. Averyanova . Yaowu Xing . 2020 . New Paleogene angiosperm species of Zaissan Depression (eastern Kazakhstan) . Botanicheskii Zhurnal . 105 . 1 . 46–57 . 10.31857/S0006813620010044 . 218793001 .
19. Xiaoyan Liu . Hanzhang Song . Jianhua Jin . 2020 . Diversity of Fagaceae on Hainan Island of South China during the middle Eocene: implications for phytogeography and paleoecology . Frontiers in Ecology and Evolution . 8 . Article 255 . 10.3389/fevo.2020.00255 . 221117336 . free .
20. Li Xue . Linbo Jia . Gi-soo Nam . Yongjiang Huang . Shitao Zhang . Yuqing Wang . Zhuo Zhou . Yongsheng Chen . 2020 . Involucre fossils of Carpinus, a northern temperate element, from the Miocene of China and the evolution of its species diversity in East Asia . Plant Diversity . 42 . 3 . 155–167. 10.1016/j.pld.2020.01.001 . 32695948 . 7361179 . free .
21. Anthony L. Swinehart . James O. Farlow . 2020 . Plant and invertebrate macrofossils from the Pipe Creek Sinkhole (Late Neogene), Grant County, Indiana . Historical Biology: An International Journal of Paleobiology . 33 . 11 . 3111–3140 . 10.1080/08912963.2020.1851686 . 230538832 .
22. Dimitra Mantzouka . Miroslav Ivanov . Vladimir Bozukov . 2020 . The first report of an 'evergreen Castanopsis type' wood (Fagaceae) for the Late Miocene–Early Pliocene of Europe (Bulgaria, Blagoevgrad Graben) . Geological Journal . 56 . 2 . 786–803 . 10.1002/gj.3919 . free .
23. Eva-Maria Sadowski . Alexander R. Schmidt . Thomas Denk . 2020 . Staminate inflorescences with in situ pollen from Eocene Baltic amber reveal high diversity in Fagaceae (oak family) . Willdenowia . 50 . 3 . 405–517 . 10.3372/wi.50.50303 . free .
24. Cédric Del Rio . Teng-Xiang Wang . Jia Liu . Shui-Qing Liang . Robert A. Spicer . Fei-Xiang Wu . Zhe-Kun Zhou . Tao Su . 2020 . Asclepiadospermum gen. nov., the earliest fossil record of Asclepiadoideae (Apocynaceae) from the early Eocene of central Qinghai-Tibetan Plateau, and its biogeographic implications . American Journal of Botany . 107 . 1 . 126–138 . 10.1002/ajb2.1418 . 31944266 . free .
25. Cédric Del Rio . Jian Huang . Gregory W. Stull . Rémi Allemand . Zhe-Kun Zhou . Tao Su . 2020 . First macrofossil record of Icacinaceae in East Asia (early Oligocene, Wenshan Basin) and its ecological implications . Journal of Systematics and Evolution . 60 . 2 . 445–455 . 10.1111/jse.12700 . 228976920 .
26. Cédric Del Rio . Dario De Franceschi . 2020 . Icacinaceae fossil fruits from three sites of the Paris Basin (early Eocene, France): local diversity and global biogeographic implications . Geodiversitas . 42 . 2 . 17–28 . 10.5252/geodiversitas2020v42a2 . free .
27. Andrew C. Rozefelds . Gregory Stull . Peta Hayes . David R. Greenwood . 2020 . The fossil record of Icacinaceae in Australia supports long-standing Palaeo-Antarctic rainforest connections in southern high latitudes . Historical Biology: An International Journal of Paleobiology . 33 . 11 . 2854–2864. 10.1080/08912963.2020.1832089 . 229005088 .
28. Gregory Stull . Bruce H. Tiffney . Steven R. Manchester . Cédric Del Rio . Scott L. Wing . 2020 . Endocarps of Pyrenacantha (Icacinaceae) from the early Oligocene of Egypt . International Journal of Plant Sciences . 181 . 4 . 432–442 . 10.1086/706854 . 208558190 .
29. Ünal Akkemik . Dimitra Mantzouka . Umut Tunç . Fikret Koçbulut . 2020 . The first paleoxylotomical evidence from the Mid-Eocene Climate Optimum from Turkey . Review of Palaeobotany and Palynology . 285 . Article 104356 . 10.1016/j.revpalbo.2020.104356 . 229395041 .
30. Daniela P. Ruiz . M. Sol Raigemborn . Mariana Brea . Roberto R. Pujana . 2020 . Paleocene Las Violetas Fossil Forest: Wood anatomy and paleoclimatology . Journal of South American Earth Sciences . 98 . Article 102414 . 10.1016/j.jsames.2019.102414 . 2020JSAES..9802414R . 213796947 .
31. Kenton L. Chambers . George O. Poinar, Jr . 2020 . Thymolepis toxandra gen. et sp. nov., a mid-Cretaceous fossil flower with horseshoe-shaped anthers . Journal of the Botanical Research Institute of Texas . 14 . 1 . 57–64 . 10.17348/jbrit.v14.i1.896 . free .
32. George O. Poinar, Jr . Kenton L. Chambers . Urszula T. Iwaniec . Fernando E. Vega . 2020 . Valviloculus pleristaminis gen. et sp. nov., a Lauralean fossil flower with valvate anthers from mid-Cretaceous Myanmar amber . Journal of the Botanical Research Institute of Texas . 14 . 2 . 359–366 . 10.17348/jbrit.v14.i2.1014 . free .
33. Jun-Ling Dong . Bai-Nian Sun . Ai-Jing Li . Hui Chen . 2020 . The diversity of Smilax (Smilacaceae) leaves from the Middle Miocene in southeastern China . Geological Journal . 56 . 2 . 744–757 . 10.1002/gj.3882 . 225790148 .
34. Lu-Liang Huang . Jian-Hua Jin . Cheng Quan . Alexei A. Oskolski . 2020 . Mummified Magnoliaceae woods from the upper Oligocene of South China, with biogeography, paleoecology, and wood trait evolution implications . Journal of Systematics and Evolution . 58 . 1 . 89–100 . 10.1111/jse.12480 . 91861708 .
35. Zlatko Kvaček . Vasilis Teodoridis . Thomas Denk . 2020 . The Pliocene flora of Frankfurt am Main, Germany: taxonomy, palaeoenvironments and biogeographic affinities . Palaeobiodiversity and Palaeoenvironments . 100 . 3 . 647–703 . 10.1007/s12549-019-00391-6 . 2020PdPe..100..647K . free .
36. Else Marie Friis . Peter R. Crane . Kaj Raunsgaard Pedersen . 2020 . Melloniflora, a new extinct multiparted flower from the Early Cretaceous of Virginia, USA . International Journal of Plant Sciences . 181 . 9 . 887–897 . 10.1086/710490 . 224837365 .
37. Atsufumi Narita . Atsushi Yabe . Kazuhiko Uemura . Midori Matsumoto . 2020 . Late middle Miocene Konan flora from northern Hokkaido, Japan . Acta Palaeobotanica . 60 . 2 . 259–295 . 10.35535/acpa-2020-0012 . free .
38. Johanna Baez . Alexandra Crisafulli . 2020 . Novelties in the xylotaphoflora from Chiquimil Formation (Miocene), Catamarca-Argentina . Journal of South American Earth Sciences . 107 . Article 102943 . 10.1016/j.jsames.2020.102943 . 225109309 .
39. Mahasin Ali Khan . Robert A. Spicer . Teresa E. V. Spicer . Kaustav Roy . Manoshi Hazra . Taposhi Hazra . Sumana Mahato . Sanchita Kumar . Subir Bera . 2020 . Dipterocarpus (Dipterocarpaceae) leaves from the K-Pg of India: a Cretaceous Gondwana presence of the Dipterocarpaceae . Plant Systematics and Evolution . 306 . 6 . Article 90 . 10.1007/s00606-020-01718-z . 2020PSyEv.306...90K . 228870254 .
40. Oris J. Rodríguez-Reyes . Emilio Estrada-Ruiz . 2020 . Two new reports of ancient rainforest trees from the Azuero Peninsula, Panama . Ameghiniana . 57 . 3 . 209–218 . 10.5710/AMGH.22.02.2020.3299 . 216250364 .
41. Maria C. Zamaloa . Maria A. Gandolfo . Kevin C. Nixon . 2020 . 52 million years old Eucalyptus flower sheds more than pollen grains . American Journal of Botany . 107 . 12 . 1763–1771 . 10.1002/ajb2.1569 . 33274448 . 7839439 . free .
42. Eliana Moya . Mariana Brea . 2020 . Combretaceous fossil wood from Ituzaingó Formation (Late Miocene?), Argentina, indicate a coastal marine environment . Review of Palaeobotany and Palynology . 281 . Article 104270 . 10.1016/j.revpalbo.2020.104270 . 2020RPaPa.28104270M . 11336/142562 . 224947789 . free .
43. Ya Li . Yi-Ming Cui . Carole T. Gee . Xiao-Qing Liang . Cheng-Sen Li . 2020 . Primotrapa gen. nov., an extinct transitional genus bridging the evolutionary gap between Lythraceae and Trapoideae, from the early Miocene of North China . BMC Evolutionary Biology . 20 . 1 . 150 . 10.1186/s12862-020-01697-2 . 33183234 . 7661254 . 2020BMCEE..20..150L . free .
44. David Peris . Conrad C. Labandeira . Eduardo Barrón . Xavier Delclòs . Jes Rust . Bo Wang . 2020 . Generalist pollen-feeding beetles during the mid-Cretaceous . iScience . 23 . 3 . Article 100913 . 10.1016/j.isci.2020.100913 . 32191877 . 7113562 . 2020iSci...23j0913P .
45. Nathan A. Jud . Maria A. Gandolfo . 2020 . Fossil evidence from South America for the diversification of Cunoniaceae by the earliest Palaeocene . Annals of Botany . 127 . 3 . 305–315 . 10.1093/aob/mcaa154 . 32860407 . 7872129 .
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47. Raymond J. Carpenter . Lynne A. Milne . 2020 . New species of xeromorphic Banksia (Proteaceae) foliage and Banksia-like pollen from the late Eocene of Western Australia . Australian Journal of Botany . 68 . 3 . 165–178 . 10.1071/BT19110 . 214113278 .
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