2018 in paleoichthyology explained

This list of fossil fish described in 2018 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fish, bony fish, and other fish of every kind that are scheduled to be described during the year 2018, as well as other significant discoveries and events related to paleontology of fish that are scheduled to occur in 2018.

Research

A study on the fossil fish occurrences and habitat during the middle Paleozoic (480 million to 360 million years ago) is published by Sallan et al. (2018), fish originated in restricted, shallow intertidal-subtidal environments.^[1]
A survey of Devonian fish fauna from Michigan is published by Stack & Sallan (2018).^[2]
A study on the ecological diversification of thelodonts is published by Ferrón et al. (2018).^[3]
The first occurrence of pelvic girdles and intromittent organs in Euphanerops longaevus, associated with a morphologically differentiated region of the axial skeleton, is reported by Chevrinais et al. (2018).^[4]
A study on the identity of the aspidin (a primitive bone-like tissue of heterostracans) is published by Keating et al. (2018), who interpret aspidin as an acellular dermal bone.^[5]
Redescription of Tesseraspis mosaica is published by Blieck, Elliott & Karatajūtė-Talimaa (2018).^[6]
A study on the morphological and taxonomic diversity of pteraspidiforms is published by Romano, Sansom & Randle (2018).^[7]
A study on the diversity of jaw shapes in modern and Paleozoic jawed fishes, evaluating whether the full extent of jaw morphological variation was established early in gnathostome evolutionary history, is published by Hill et al. (2018).^[8]
New specimens of Brindabellaspis stensioi, providing new information on the morphology of the rostral region of the skull, are described from the Lower Devonian of the New South Wales (Australia) by King, Young & Long (2018).^[9]
Redescription of the antiarch placoderm Asterolepis thule and a study on the age of the deposits preserving the fossils of this species is published by Newman & Den Blaauwen (2018).^[10]
Description of bony pelvic plates in 32 specimens of Bothriolepis canadensis from the Upper Devonian Escuminac Formation (Canada) is published by Charest, Johanson & Cloutier (2018), who reject the interpretation of these structures as genital plates (suggested by Long et al., 2015),^[11] and identify them as the pelvic girdle instead.^[12]
Redescription of the antiarch placoderm Phymolepis cuifengshanensis and a study on the phylogenetic relationships of this species is published by Wang & Zhu (2018).^[13]
A study on the morphology of the skull, especially the braincase of the petalichthyid placoderm Shearsbyaspis oepiki is published by Castiello & Brazeau (2018).^[14]
Fossil interpreted as placoderm (arthrodiran) egg cases are described from the Devonian (Famennian) Cleveland Shale (Ohio, United States) by Carr & Jackson (2018).^[15]
Circular or near-circular patterned trace fossils, similar to underwater circles produced by male pufferfishes, are described from the Upper Devonian Hongguleleng Formation (China) by Zong & Gong (2018), who consider it possible that these fossils may be structures made by male fish to attract females.^[16]
Redescription of Gladbachus adentatus and a study on the phylogenetic relationships of the species is published by Coates et al. (2018).^[17]
A study on the wear of a tooth whorl of a specimen of Edestus heinrichi, as well as on its implications for inferring the function of the tooth whorls in this species, is published by Itano (2018).^[18]
New description of Edestus, providing new information on the anatomy of this taxon, is published online by Tapanila et al. (2018).^[19]
Two partial specimens of a callorhynchid chimaeroid left in open nomenclature are described from the Upper Kimmeridgian Nusplingen Plattenkalk (Germany) by Duffin (2018).^[20]
A study on the teeth histology and vasculature of the oldest known tooth-bearing sharks, Leonodus carlsi and Celtiberina maderi, is published by Martinez-Perez et al. (2018).^[21]
The first known basicranium of Carcharopsis wortheni is described from the Carboniferous Fayetteville Shale (United States) by Bronson, Mapes & Maisey (2018).^[22]
A study on the morphology of the braincase of Tristychius arcuatus is published by Coates & Tietjen (2018).^[23]
A diverse fauna of Early Triassic cartilaginous fishes is described from the Vikinghøgda Formation (Spitsbergen, Norway) by Bratvold, Delsett & Hurum (2018).^[24]
The first material referable to hybodont shark (a member of the genus Asteracanthus) is described from the Lower Jurassic (Toarcian) Rosso Ammonitico Formation (Italy) by Romano et al. (2018), providing new information on the dispersal of this genus in the Jurassic Tethys.^[25]
The first scroll coprolites from the Mesozoic reported so far, likely produced by euryhaline hybodontid sharks, are described from the Upper Triassic Tiki Formation of India by Rakshit et al. (2018), who name a new coprolite taxon Tikicopros triassicus.^[26]
Description of new remains of the Late Jurassic shark Palaeocarcharias stromeri and a study on the anatomy and phylogenetic relationships of this species is published by Landemaine, Thies & Waschkewitz (2018), who name a new order Palaeocarchariiformes and a new family Palaeocarchariidae.^[27]
Description of an articulated skeleton of a member of the palaeospinacid genus Synechodus from the Lower Cretaceous (Albian) Saint-Pô Formation (France), and a revision of the taxonomic history of the species assigned to the genus Synechodus, is published by Mollen & Hovestadt (2018).^[28]
Late Cretaceous taxon Platylithophycus cretaceus known from the Niobrara Chalk of Kansas (United States), considered to be a green alga or a cuttlefish in earlier publications, is reinterpreted as a member of Elasmobranchii by Bronson & Maisey (2018).^[29]
A study on the microstructure of enameloid in the isolated teeth of archaeobatid batomorphs Toarcibatis elongata, Cristabatis crescentiformis and Doliobatis weisi from the Jurassic (Toarcian) localities of Halanzy (Belgium) and Ginzebierg (Luxembourg) is published by Manzanares, Botella & Delsate (2018).^[30]
A study on the structure of teeth of Myledaphus pustulosus from the Upper Cretaceous (Maastrichtian) Hell Creek Formation (Montana, United States) is published by Hoffman, Jensen & Hageman (2018).^[31]
Isolated teeth of the sand shark Brachycarcharias lerichei are described from the Eocene (Ypresian) La Meseta Formation (Antarctica) by Marramà et al. (2018), representing the southernmost occurrence of the genus Brachycarcharias reported so far.^[32]
A study on the anatomy, paleobiology and paleoecology of the Eocene requiem shark Eogaleus bolcensis is published by Marramà, Carnevale & Kriwet (2018).^[33]
Teeth of members of the genera Galeorhinus and Physogaleus are described from the Lower Eocene sediments of the Khuiala Formation (Jaisalmer basin, India) by Pandey, Chaskar & Case (2018).^[34]
A study on the teeth mineralization process and teeth histology in extant and fossil members of the genus Hemipristis is published by Jambura et al. (2018).^[35]
A study on the global and regional morphological variation of the teeth of the ground sharks and mackerel sharks across the Cretaceous–Paleogene boundary is published by Bazzi et al. (2018).^[36]
A study on the anatomy and evolution of teeth of members of the families Megachasmidae and Cetorhinidae, based on data from recent and fossil teeth, is published by Mitchell, Ciampaglio & Jacquemin (2018).^[37]
A study on the physiological, ecological and life-history traits which influenced the biogeographic distributions of cartilaginous fishes from the Neogene to the present is published by Villafaña & Rivadeneira (2018).^[38]
A review of the present and past (Miocene–Pleistocene) shark and ray diversity in marine waters of Tropical America is published by Carrillo-Briceño et al. (2018).^[39]
A study on the phylogenetic relationships of extant and fossil squalomorph sharks as indicated by teeth morphology is published by Flammensbeck et al. (2018).^[40]
A study on the morphology and phylogenetic relationships of an early bony fish known from two partial skulls recovered from the Devonian (Emsian) Taemas Limestones of the Burrinjuck area (New South Wales, Australia), possibly belonging to the genus Ligulalepis (described on the basis of isolated scales), is published by Clement et al. (2018).^[41]
Redescription of Elonichthys germari is published by Schindler (2018), who presents the first reconstruction of the skull of this species.^[42]
Redescription of the neotype of the elonichthyid Rhabdolepis macropterus is published by Schindler (2018), who presents new reconstruction of the skull of this species.^[43]
A revision of ray-finned fishes from the Permian locality Buxières-les-Mines (Bourbon-l’Archambault Basin, France) is published by Štamberg (2018).^[44]
A study on the effect of Permian–Triassic and Triassic–Jurassic extinction events on ray-finned fishes is published by Smithwick & Stubbs (2018).^[45]
A study on the evolutionary history of ray-finned fishes across the Cretaceous–Paleogene extinction event, as indicated by isolated fossil teeth preserved in a South Pacific sediment core spanning 72–43 Ma, is published by Sibert et al. (2018).^[46]
A study on the morphological variation of the dorsal finlets in extant bichirs, testing the viability of these anatomic structures as a tool for taxonomic diagnoses in the study of fossil members of this group, is published by Coelho, Cupello & Brito (2018).^[47]
New data on the reproductive biology of the species Saurichthys curionii and Saurichthys macrocephalus from the Middle Triassic Meride Limestone (Monte San Giorgio, Switzerland) is presented by Maxwell et al. (2018), who identify six specimens as unambiguously gravid.^[48]
A study on the internal anatomy of the skulls of two Early Triassic specimens of Saurichthys, as well as on the phylogenetic relationships of saurichthyiforms, is published by Argyriou et al. (2018).^[49]
A comparative study on the bony labyrinth of early neopterygians, including relatives of gars and teleosts, is published by Giles, Rogers & Friedman (2018).^[50]
A study on the diversity of body shapes of neopterygians from the Triassic to the Early Cretaceous is published by Clarke & Friedman (2018).^[51]
Redescription and taxonomic reassessment of the pycnodontiform genus Cosmodus is published by Vullo et al. (2018).^[52]
A study on 52 specimens of Pycnodus from the Eocene Monte Bolca Lagerstätte (Italy), evaluating whether the morphological variability within the sample might be related to inter- or intraspecific variation, is published by Cawley et al. (2018).^[53]
A study on the anatomical structure and possible function of the flank bar-scales of members of Pycnodontiformes is published by Capasso (2018).^[54]
A revision of the phylogenetic relationships of the fossils fishes belonging to the group Halecomorphi is published by Ebert (2018).^[55]
A redescription of Asialepidotus shingyiensis and a study on the phylogenetic relationships of the species is published by Xu & Ma (2018).^[56]
A study on the phylogenetic relationships of the Triassic neopterygian Ticinolepis, as well as on the relationships of the fossil neopterygians in general, is published by López-Arbarello & Sferco (2018).^[57]
A study on the locomotion energetics of Leedsichthys problematicus, possible factors that drove the gigantism in pachycormiforms and the metabolic limits of body size in ray-finned fishes is published by Ferrón et al. (2018).^[58]
An ichthyodectiform fossil specimen preserving a small skull and anterior part of the trunk is described from a core recovered from a well drilled in the Cape Verde Basin, ca. 400 km offshore from the West African Atlantic Margin, by Casson et al. (2018).^[59]
A study on the evolutionary history of the family Catostomidae, based on data from molecules, morphology and fossil record, is published by Bagley, Mayden & Harris (2018).^[60]
A study on the phylogenetic relationships of members of Acanthomorpha and on the timescale of the radiation of this group is published by Alfaro et al. (2018), who report that crown ages for five of the six major percomorph subclades, and for the bulk of the species diversity in the sixth, coincide with the Cretaceous–Paleogene boundary.^[61]
A study on the morphology of the skeleton of Pholidophorus latiusculus, as well as on the phylogenetic relationships of this species, is published by Taverne (2018).^[62]
A study on the gill-arch anatomy in Late Cretaceous–early Paleogene members of Aulopiformes is published by Beckett, Giles & Friedman (2018).^[63]
A new specimen of Spinocaudichthys with preserved intestinal tract is described from the Cretaceous (Cenomanian) Jbel Oum Tkout Lagerstätte (Morocco) by Davesne et al. (2018).^[64]
A study on the bone histology of extant opahs, comparing it with bones of their extant and fossil relatives (including "Aipichthys" velifer), and testing the hypothesized link between endothermy and cellular bone (bone containing embedded osteocyte cells) in teleosts, is published by Davesne et al. (2018).^[65]
An articulated skeleton of a juvenile toadfish distinct from Louckaichthys novosadi is described from the Oligocene Bituminous Marls Formation (Romania) by Pikryl et al. (2018).^[66]
A study on the dynamics of diversification, phenotypic evolution and habitat transitions in the ray-finned fish group Carangaria after the Cretaceous–Paleogene extinction event is published by Ribeiro et al. (2018).^[67]
A study on the phylogenetic relationships of the fossil snake mackerels and cutlassfishes from the Eocene (Ypresian) London Clay Formation (United Kingdom) is published by Beckett et al. (2018).^[68]
Detailed description of the caudal skeleton of the Paleogene surgeonfish Arambourgthurus scombrurus is published by Carnevale & Tyler (2018).^[69]
A study on the morphology of the Oligocene percoid fish Oligoserranoides budensis is published by Bieńkowska-Wasiluk & Pałdyna (2018).^[70]
A study on the Pliocene fish fossils from the Kanapoi site (Kenya) and their implications for reconstructing lake and river environments in the Kanapoi Formation is published online by Stewart & Rufolo (2018).^[71]
A metacarpal bone of a specimen of Pteranodon, bearing teeth marks likely produced by a shark and by a saurodontid fish, is described from the Campanian Mooreville Chalk (Alabama, United States) by Ehret & Harrell (2018).^[72]
A series of neck vertebrae of Pteranodon associated with a tooth of the lamniform shark Cretoxyrhina mantelli is described from the Upper Cretaceous Niobrara Formation (Kansas, United States) by Hone, Witton & Habib (2018), who interpret the specimen as evidence of Cretoxyrhina biting Pteranodon.^[73]
A mawsoniid coelacanth specimen is described from Rhaetian deposits of the Var Department (France) by Deesri et al. (2018), representing the first known coelacanth from the marine Triassic of France.^[74]
A study on both newly collected and earlier fossil material of Ventalepis ketleriensis from the Devonian (Famennian) of Latvia and central and northwestern Russia is published by Lebedev & Lukševičs (2018), who interpret the fossils as supporting the porolepiform affinities of this species, and name a new family Ventalepididae.^[75]
Anatomical description of the endocast of "Chirodipterus" australis from the Upper Devonian Gogo Formation (Australia) is published by Henderson & Challands (2018).^[76]
A revision of the lungfish remains from the Triassic of the Świętokrzyskie Mountains and from the northeastern Poland is published by Skrzycki, Niedźwiedzki & Tałanda (2018), who report the first known Middle Triassic finding of Arganodus worldwide and the oldest known occurrence of Ptychoceratodus in Europe.^[77]
Lungfish burrows are reported for the first time from the Lower and Middle Triassic deposits of the Southern Cis-Urals by Sennikov (2018).^[78]
A study on the anatomy of the lungfish Mioceratodus gregoryi from the Eocene Redbank Plains Formation (Australia) is published by Kemp (2018).^[79]
Description of well-preserved pelvic fin skeleton of a specimen of Rhizodus hibberti from the Carboniferous (Viséan) Asbian Wardie Shales (United Kingdom) is published by Jeffery et al. (2018).^[80]
A rediagnosis and redescription of Hyneria lindae based on new remains from the Catskill Formation (Pennsylvania, United States) is published by Daeschler & Downs (2018).^[81]

New taxa

Jawless vertebrates

Name	Novelty	Status	Authors	Age	Unit	Notes
Altigibbaspis^[82]	Gen. et sp. nov	Valid	Liu, Gai & Zhu	Devonian (early Lochkovian)	Xishancun	A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Polybranchiaspidae. The type species is A. huiqingae.
Elgaia^[83]	Gen. et comb. nov	Valid	Glinskiy	Devonian		A member of the family Psammosteidae. The type species is "Tartuosteus" luhai Mark-Kurik (1965).
Faberaspis^[84]	Gen. et sp. nov	Valid	Elliott, Lassiter & Blieck	Devonian (Lochkovian)	Drake Bay	A member of the family Cyathaspididae. The type species is F. elgae.
Kalanaspis^[85]	Gen. et sp. nov	Valid	Tinn & Märss	Silurian (Aeronian)		An early osteostracan. Genus includes new species K. delectabilis.
Nanningaspis^[86]	Gen. et sp. nov	Valid	Gai et al.	Devonian (Pragian)	Nakaolin	A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Gumuaspidae. The type species is N. zengi.
Paralogania denisoni^[87]	Sp. nov	Valid	Turner & Burrow	Silurian–Devonian boundary	Eastport	A thelodont.
Platylomaspis	Gen. et sp. nov	Valid	Gai et al.	Silurian (Telychian)	Tataertag	A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Gumuaspidae. The type species is P. serratus.
Pseudolaxaspis	Gen. et comb. nov	Valid	Gai et al.	Devonian (early Lochkovian)	Tataertag	A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Gumuaspidae. The type species is "Laxaspis" rostrata Liu (1975).
Vladimirolepis	Gen. et comb. nov	Valid	Glinskiy	Devonian		A member of the family Psammosteidae. The type species is "Psammolepis" proia Mark-Kurik (1965).

Placoderms

Name	Novelty	Status	Authors	Age	Unit	Notes
Hlavinichthys^[88]	Gen. et sp. nov	Valid	Carr	Devonian (Famennian)	Ohio Shale	A member of Aspinothoracidi. The type species is H. jacksoni.
Microbrachius kedoae^[89]	Sp. nov	Valid	Mark-Kurik et al.	Devonian (Givetian)	Moroch Beds
Wufengshania^[90]	Gen. et sp. nov	Valid	Pan et al.	Devonian (Emsian)		A member of Antiarchi belonging to the family Bothriolepididae. The type species is W. magniforaminis.

Acanthodians

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Cheiracanthus flabellicostatus^[91]	Sp. nov	Valid	Pinakhina	Middle Devonian		^[92]
Drygantacanthus^[93]	Gen. et sp. nov	Valid	Voichyshyn & Szaniawski	Early Devonian			A member of Ischnacanthiformes belonging to the new family Podoliacanthidae. The type species is D. semirotunda.
Ginkgolepis^[94]	Gen. et sp. et comb. nov	Valid	Pinakhina & Märss	Devonian (Eifelian, Givetian, probably Famennian)	Härma Beds		A member of Acanthodiformes belonging to the family Cheiracanthidae. The type species is G. tenericostatus; genus also includes "Cheiracanthus" talimae Valiukevičius (1985).
Jolepis^[95]	Gen. et comb. nov	Valid	Burrow & Turner	Devonian (Lochkovian)			A possible acanthodian of uncertain phylogenetic placement. The type species is "Diplacanthoides" robustus Brotzen (1934).
Kasperacanthus	Gen. et sp. nov	Valid	Voichyshyn & Szaniawski	Early Devonian			A member of Ischnacanthiformes belonging to the new family Podoliacanthidae. The type species is K. serratus.
Rhadinacanthus deltosquamosus	Sp. nov	Valid	Pinakhina & Märss	Devonian (Givetian)	Härma Beds		A member of Diplacanthiformes belonging to the family Diplacanthidae.

Cartilaginous fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acutalamna^[96]	Gen. et sp. nov	Valid	Guinot & Carrillo-Briceño	Cretaceous (Albian and Cenomanian)	La Luna		A mackerel shark of uncertain phylogenetic placement. Genus includes new species A. karsteni.
Alopias palatasi^[97]	Sp. nov	Valid	Kent & Ward	Miocene	Calvert	( )	A thresher shark.
Altholepis salopensis	Sp. nov	Valid	Burrow & Turner	Devonian (Lochkovian)			A member of Altholepidiformes belonging to the family Altholepididae.
Altusmirus^[98]	Gen. et sp. nov	Valid	Fuchs et al.	Early Cretaceous (Valanginian)			A ground shark. Genus includes new species A. triquetrus.
Amaradontus	Gen. et sp. nov	Valid	Hodnett & Elliott	Carboniferous (Serpukhovian)	Surprise Canyon		A member of Neoselachii belonging to the family Anachronistidae. The type species is A. santuccii.
Arcuodus^[99]	Gen. et sp. et comb. nov	Valid	Itano & Lambert	Carboniferous (late Tournaisian to Viséan)	Bangor Burlington Keokuk Salem	( )	A member of Holocephali belonging to the group Cochliodontiformes. The type species is A. multicuspidatus; genus also includes "Deltodopsis" bialveatus St. John & Worthen (1883).
Carcharoides lipsiensis^[100]	Sp. nov	Valid	Reinecke et al.	Oligocene (Rupelian)	Böhlen		A sand shark.
Caucasochasma^[101]	Gen. et sp. nov	Valid	Prokofiev & Sychevskaya	Early Oligocene			A relative of the basking shark. The type species is C. zherikhini.
Cooleyella platera	Sp. nov	Valid	Hodnett & Elliott	Carboniferous (Serpukhovian)	Surprise Canyon		A member of Neoselachii belonging to the family Anachronistidae.
Cretacladoides^[102]	Gen. et 2 sp. nov	Valid	Feichtinger et al.	Early Cretaceous (Valanginian)			A cartilaginous fish of uncertain affinities. Originally described as a possible member of the family Falcatidae; Ivanov (2022) considered its teeth to be only superficially similar to the teeth of Paleozoic falcatids, and considered it to be a possible neoselachian.^[103] Genus includes new species C. ogiveformis and C. noricum.
Cretalamna bryanti^[104]	Sp. nov	Valid	Ebersole & Ehret	Late Cretaceous (late Santonian and early Campanian)	Eutaw Mooreville Chalk
Deltodus tubineus^[105]	Sp. nov	Valid	Richards et al.	Carboniferous (Tournaisian)	Ballagan		A member of Cochliodontiformes belonging to the family Cochliodontidae.
Fornicatus	Gen. et sp. nov	Valid	Fuchs et al.	Early Cretaceous (Valanginian)			A ground shark. Genus includes new species F. austriacus.
Hokomata^[106]	Gen. et sp. nov	Valid	Hodnett & Elliott	Carboniferous (Bashkirian)	Watahomigi		A member of Xenacanthiformes belonging to the family Diplodoselachidae. The type species is H. parva.
Hybodus chuanjieensis^[107]	Sp. nov		Sun in Sun et al.	Middle Jurassic	Chuanjie Basin
Hybodus xinzhuangensis	Sp. nov		Sun in Sun et al.	Middle Jurassic	Chuanjie Basin
Igdabatis marmii^[108]	Sp. nov	Valid	Blanco	Late Cretaceous (Maastrichtian)
Lamarodus^[109]	Gen. et sp. nov	Valid	Ivanov in Ivanov et al.	Permian (Guadalupian)	Bell Canyon		A member of Hybodontiformes belonging to the superfamily Hybodontoidea. Genus includes new species L. triangulus. Announced in 2018; the final version of the article naming it was published in 2020.
Marambioraja^[110]	Gen. et sp. nov	Valid	Engelbrecht et al.	Eocene (Ypresian)	La Meseta	Antarctica (Seymour Island)	A skate. The type species is M. leiostemma.
Mesetaraja	Gen. et sp. nov	Valid	Engelbrecht et al.	Eocene (Ypresian)	La Meseta	Antarctica (Seymour Island)	A skate. The type species is M. maleficapelli.
Microcarcharias	Gen. et comb. nov	Valid	Guinot & Carrillo-Briceño	Late Cretaceous	Britton Niobrara		A sand shark; a new genus for "Odontaspis" saskatchewanensis Case, Tokaryk & Baird (1990).
Microklomax	Gen. et sp. nov	Valid	Hodnett & Elliott	Carboniferous (Serpukhovian)	Surprise Canyon		A member of Euselachii belonging to the family Protacrodontidae. The type species is M. carrieae.
Mooreodontus jaini^[111]	Sp. nov	Valid	Bhat, Ray & Datta	Late Triassic	Tiki		A xenacanthid.
Natarapax	Gen. et sp. nov	Valid	Feichtinger et al.	Early Cretaceous (Valanginian)			A possible member of the family Ctenacanthidae. Genus includes new species N. trivortex.
Novaculodus	Gen. et sp. nov	Valid	Hodnett & Elliott	Carboniferous (Serpukhovian)	Surprise Canyon		A member of Euselachii belonging to the family Protacrodontidae. The type species is N. billingsleyi.
Ostarriraja^[112]	Gen. et sp. nov	Valid	Marramà, Schultz & Kriwet	Miocene (Burdigalian)			A member of Rajiformes of uncertain phylogenetic placement. The type species is O. parva.
Palaeocentroscymnus^[113]	Gen. et comb. nov	Valid	Pollerspöck, Flammensbeck & Straube	Miocene	Lakšárska Nová Ves		A sleeper shark; a new genus for "Paraetmopterus" horvathi Underwood & Schlögl (2013).
Palaeoheterodontus^[114]	Gen. et comb. nov	Valid	Hovestadt	Jurassic			A bullhead shark. Genus includes "Heterodontus" sarstedtensis Thies (1983).
Pastinachus kebarensis^[115]	Sp. nov	Valid	Adnet et al.	Eocene (late Bartonian)			A cowtail stingray of Pastinachus. Announced in 2018; the final version of the article naming it was published in 2019.
Pristrisodus^[116]	Gen. et comb. nov	Valid	Bhat, Ray & Datta	Late Triassic	Tiki		A member of Hybodontiformes belonging to the family Lonchidiidae. Genus includes "Parvodus" tikiensis Prasad et al. (2008).
Procestracion	Gen. et comb. nov	Valid	Hovestadt	Jurassic			A bullhead shark. Genus includes "Acrodus" semirugosus Plieninger (1847), "Hemipristis" bidens Quenstedt (1852) and "Strophodus" semirugosus Quenstedt (1852).
Protoheterodontus	Gen. et comb. nov	Valid	Hovestadt	Late Cretaceous			A bullhead shark. Genus includes "Heterodontus" boussioni Guinot et al. (2013).
Protohimantura^[117]	Gen. et comb. nov	Valid	Marramà et al.	Miocene (Burdigalian?)	Tonasa		A whiptail stingray belonging to the subfamily Urogymninae. The type species is "Trygon" vorstmani de Beaufort (1926).
Raja amphitrita	Sp. nov	Valid	Engelbrecht et al.	Eocene (Ypresian and Lutetian)	La Meseta Submeseta	Antarctica (Seymour Island)	A skate, a species of Raja.
Raja manitaria	Sp. nov	Valid	Engelbrecht et al.	Eocene (Ypresian and Lutetian)	La Meseta Submeseta	Antarctica (Seymour Island)	A skate, a species of Raja.
Similiteroscyllium	Gen. et sp. et comb. nov	Valid	Fuchs et al.	Early Cretaceous (Valanginian)			A carpet shark. Genus includes new species S. iniquus, as well as "Ornatoscyllium" rugasimulatum Guinot, Cappetta & Adnet (2014).
Squalicorax acutus^[118]	Sp. nov	Valid	Siversson et al.	Late Cretaceous (Cenomanian)	Gearle
Squalicorax bazzii	Sp. nov	Valid	Siversson et al.	Late Cretaceous (Cenomanian)	Gearle
Squalicorax lalunaensis	Sp. nov	Valid	Guinot & Carrillo-Briceño	Late Cretaceous (Cenomanian)	La Luna
Squalicorax moodyi	Sp. nov	Valid	Guinot & Carrillo-Briceño	Late Cretaceous (Cenomanian)	La Luna
Squalicorax mutabilis	Sp. nov	Valid	Siversson et al.	Late Cretaceous (Cenomanian and Turonian)	Haycock
Stethacanthus concavus^[119]	Sp. nov	Valid	Ginter	Carboniferous (Gzhelian)	Indian Cave
Tamiobatis elgae^[120]	Sp. nov	Valid	Ivanov in Ivanov & Plax	Devonian (Famennian) and Carboniferous (Tournaisian)			A member of Ctenacanthiformes belonging to the family Ctenacanthidae.
Tikiodontus	Gen. et sp. nov	Valid	Bhat, Ray & Datta	Late Triassic	Tiki		A xenacanthid. Genus includes new species T. asymmetricus.
Truyolsodontos^[121]	Gen. et sp. nov	Valid	Bernárdez	Late Cretaceous (Cenomanian)			A mackerel shark. Genus includes new species T. estauni.
Whitropus	Gen. et sp. nov	Valid	Richards et al.	Carboniferous (Tournaisian)	Ballagan		A member of Cochliodontiformes of uncertain phylogenetic placement. The type species is W. longicalcus.

Ray-finned fishes

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Allogenartina^[122]	Gen. et sp. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian and Campanian)	Eutaw	( )	Possibly a member of Stomiiformes belonging to the group Gonostomatoidei, of uncertain phylogenetic placement within the latter group. The type species is A. muscogeei.
Amakusaichthys^[123]	Gen. et sp. nov	Valid	Yabumoto, Hirose & Brito	Late Cretaceous (Santonian)	Hinoshima		A member of Ichthyodectiformes. Genus includes new species A. goshouraensis. Announced in 2018; the final version of the article naming it was published in 2020.
Ameiurus grangerensis^[124]	Sp. nov	Valid	Smith, Martin & Carpenter	Miocene	Ellensburg		A species of Ameiurus.
Amphiarius paleoorinocoensis^[125]	Sp. nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco		A species of Amphiarius.
Anchiacipenser^[126]	Gen. et sp. nov	Valid	Sato et al.	Late Cretaceous (Campanian)	Dinosaur Park		A sturgeon. Genus includes new species A. acanthaspis.
Andrewsolepis^[127]	Gen. et sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Haplolepidae. The type species is A. lochlani.
Apateodus? assisi	Sp. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian)	Eutaw		A member of Aulopiformes belonging to the family Ichthyotringidae.
Aphanolebias sarmaticus^[128]	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene
Apsopelix ?berlinensis^[129]	Sp. nov	Valid	Schwarzhans	Early Cretaceous (Berriasian to Valanginian)			A member of Crossognathiformes belonging to the family Crossognathidae.
Archaeglossus	Gen. et comb. et sp. nov	Valid	Schwarzhans	Jurassic (Bathonian to Tithonian)			A member of Osteoglossiformes of uncertain phylogenetic placement. The type species is "Otolithus (Leptolepidarum)" pentangulatus Frost (1924); genus also includes new species A. torrensi.
Ariopsis ariopsilus	Sp. nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco		A species of Ariopsis.
Atherina carnevalei	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene			A species of Atherina.
Aulothrissus	Gen. et 2 sp. nov	Valid	Schwarzhans	Early Cretaceous (Berriasian to Valanginian)	Gevaram		A member of Elopiformes of uncertain phylogenetic placement. The type species is A. avitus; genus also includes A. heletzensis.
Avonichthys^[130]	Gen. et sp. nov		Wilson, Pardo & Anderson	Carboniferous (Tournaisian)	Horton Bluff		An early ray-finned fish. The type species is A. manskyi.
Bagre urumacoensis	Sp. nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco		A species of Bagre.
Benthalbella praecessor^[131]	Sp. nov	Valid	Nazarkin & Carnevale	Miocene (late Langhian–early Serravallian)	Kurasi		A species of Benthalbella.
Bicavolithus	Gen. et sp. nov	Valid	Schwarzhans	Early Cretaceous (Albian)			A teleost of uncertain phylogenetic placement. Genus includes new species B. cavatus.
Blairolepis wallacei	Sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Haplolepidae.
Blennius? martinii	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene			A combtooth blenny.
Bluefieldius^[132]	Gen. et sp. nov	Valid	Mickle	Carboniferous (late Mississippian)	Bluefield		An early ray-finned fish. The type species is B. mercerensis.
Braccohaplolepis	Gen. et sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Haplolepidae. The type species is B. fenestratum.
Cerinichthys	Gen. et sp. nov	Valid	Ebert	Late Jurassic			A member of Halecomorphi belonging to the new order Ophiopsiformes. Genus includes new species C. koelblae.
Champsodon tethensis^[133]	Sp. nov	Valid	Bannikov	Eocene (Bartonian)			A species of Champsodon
Chaoia^[134]	Gen. et comb. nov	Valid	Bannikov, Schwarzhans & Carnevale	Neogene			A member of the family Sciaenidae. The type species is "Sciaena" moguntiniformis Pana (1977).
Clupea macrocephala^[135]	Sp. nov	Junior homonym	Yabumoto & Nazarkin	Miocene	Bessho		A species of Clupea. The specific name is preoccupied by Clupea macrocephala Lacépède (1803); Yabumoto & Nazarkin (2020) coined a replacement name Clupea hanishinaensis.^[136]
Coryphaenoides scrupus^[137]	Sp. nov	Valid	Brzobohatý & Nolf	Middle Miocene			A species of Coryphaenoides.
Cowetaichthys	Gen. et 2 sp. et comb. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian to Maastrichtian) and Paleocene	Eutaw		A beardfish. The type species is C. alabamae; genus also includes new species C. amberi, as well as "genus Polymixiidarum" beaury Schwarzhans (2010), "genus Veliferidarum" groenlandicus Schwarzhans (2004) and "genus Veliferidarum" harderi Schwarzhans (2003).
Cretolepis^[138]	Gen. et sp. nov	Valid	Wang et al.	Early Cretaceous	Longjiang		A member of Palaeonisciformes. The type species is C. dongbeiensis.
Croatosciaena	Gen. et sp. nov	Valid	Bannikov, Schwarzhans & Carnevale	Middle Miocene			A member of the family Sciaenidae. The type species is C. krambergeri.
Cubariomma^[139]	Gen. et sp. nov	Valid	Bannikov	Early Oligocene			A member of Stromateoidei of uncertain phylogenetic placement. Genus includes new species C. yanakuzminae.
Dapedium ballei^[140]	Sp. nov	Valid	Maxwell & López-Arbarello	Middle Jurassic (Aalenian)	Opalinuston
Doggerichthys	Gen. et sp. nov	Valid	Schwarzhans	Middle Jurassic (Aalenian			A member of Leptolepidiformes sensu lato of uncertain phylogenetic placement. Genus includes new species D. anguilliformis.
Eleogobius prochazkai	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene
Elops eutawanus	Sp. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian)	Eutaw		A species of Elops.
Eoalosa^[141]	Gen. et sp. nov	Valid	Marramà & Carnevale	Eocene (Ypresian)	Monte Bolca		A member of the family Clupeidae. The type species is E. janvieri.
Eoellimmichthys^[142]	Gen. et sp. nov	Valid	Marramà et al.	Eocene (late Ypresian)	Monte Bolca		A member of Clupeomorpha belonging to the group Ellimmichthyiformes and the family Paraclupeidae. The type species is E. superstes.
Eutawichthys	Gen. et 2 sp. et comb. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian to Maastrichtian)	Eutaw	( )	A member of Beryciformes belonging to the group Berycoidei, of uncertain phylogenetic placement within the latter group. The type species is E. stringeri; genus also includes new species E. compressus, as well as “genus Apogonidarum” maastrichtiensis Nolf & Stringer (1996) and “genus Apogonidarum” zideki Nolf & Stringer (1996).
Furloichthys^[143]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)			A member of the family Ichthyodectidae. The type species is F. bonarellii.
Fuyuanichthys^[144]	Gen. et sp. nov	Valid	Xu, Ma & Ren	Middle Triassic (Ladinian)	Falang		An early member of Ginglymodi. The type species is F. wangi.
Genartina cretacea	Sp. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian)	Eutaw		Possibly a member of Stomiiformes.
Gobius apuseni	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene			A species of Gobius.
Gobius holcovae	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene			A species of Gobius.
Gobius manfredi	Sp. nov	Valid	Reichenbacher, Filipescu & Miclea	Middle Miocene			A species of Gobius.
Grimmenichthys^[145]	Gen. et sp. nov	Valid	Konwert & Hörnig	Early Jurassic (Toarcian)			A member of Pholidophoriformes. Genus includes new species G. ansorgei.
Guimarotaichthys	Gen. et sp. nov	Valid	Schwarzhans	Late Jurassic	Camadas de Guimarota		A non-teleost bony fish of uncertain phylogenetic placement. Genus includes new species G. problematicus.
Haqelpycnodus^[146]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)			A member of the family Pycnodontidae. The type species is H. picteti.
Ijimaia rara	Sp. nov	Valid	Brzobohatý & Nolf	Middle Miocene			A species of Ijimaia.
Italoalbula^[147]	Gen. et sp. nov	Valid	Taverne & Capasso	Early Cretaceous (Albian)	Pietraroja		A bonefish. The type species is I. pietrarojae.
Joffrichthys tanyourus^[148]	Sp. nov	Valid	Murray et al.	Paleocene	Paskapoo		A member of Osteoglossomorpha of uncertain phylogenetic placement.
Kalabisia^[149]	Gen. et sp. nov	Valid	Přikryl & Carnevale	Oligocene–early Miocene	Ždánicko-Hustopeče		A viviparous brotula belonging to the subfamily Brosmophycinae. The type species is K. krumvirensis.
Lagocephalus striatus^[150]	Sp. nov	Valid	Aguilera et al.	Miocene (Serravallian)	Valiente Peninsula		A species of Lagocephalus.
Lanarkichthys	Gen. et sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri of uncertain phylogenetic placement. The type species is L. gardineri.
Landinisciaena	Gen. et sp. et comb. nov	Valid	Bannikov, Schwarzhans & Carnevale	Miocene (late Burdigalian or early Langhian)			A member of the family Sciaenidae. The type species is L. popovi; genus also includes "Atractoscion" elongatissimus Schwarzhans (1993).
Lanxangichthys^[151]	Gen. et sp. nov	Valid	Cavin et al.	Early Cretaceous (Aptian)	Grès supérieurs Khok Kruat?	?	A gar. Genus includes new species L. alticephalus.
Lasalichthys otischalkensis^[152]	Sp. nov	Valid	Gibson	Late Triassic	Dockum Group		A member of Redfieldiiformes.
Leptoelops	Gen. et comb. nov	Valid	Schwarzhans	Middle Jurassic			A member of Elopiformes of uncertain phylogenetic placement. The type species is Otolithus (Lycopteridarum) rhenanus Weiler (1954); genus also includes Otolithus (Lycopteridarum) acutus Weiler (1954).
Leptosciaena	Gen. et comb. et sp. nov	Valid	Bannikov, Schwarzhans & Carnevale	Neogene			A member of the family Sciaenidae. The type species is "Genyonemus" karagiensis Bratishko, Schwarzhans & Reichenbacher (2015); genus also includes "Otolithus (Mugilidarum)" azerbaidjanicus Djafarova (2006), as well as new species L. caputoi.
Leuroglossus kobylianskyi^[153] ^[154]	Sp. nov	Valid	Nazarkin	Middle–Late Miocene	Kurasi		A species of Leuroglossus
Libanopycnodus^[155]	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)			A member of the family Pycnodontidae. The type species is L. wenzi.
Lopadichthys	Gen. et sp. nov	Valid	Murray et al.	Paleocene	Paskapoo		A member of Osteoglossiformes. The type species is L. colwellae.
Macroynis	Gen. et sp. nov	Valid	Beckett et al.	Eocene	London Clay		A relative of snake mackerels and cutlassfishes. Genus includes new species M. casieri.
Motlayoichthys^[156]	Gen. et sp. nov	Valid	Arratia, González-Rodríguez & Hernández-Guerrero	Cretaceous (Albian-Cenomanian)	El Doctor		A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is M. sergioi.
Mylocheilus kevinmeeksi	Sp. nov	Valid	Smith, Martin & Carpenter	Miocene	Ellensburg		A relative of the peamouth.
Nardoglossus^[157]	Gen. et sp. nov.	Valid	Taverne & Capasso	Late Cretaceous (Campanian-Maastrichtian)	Altamura		A member of Gonorynchiformes belonging to the group Gonorynchoidei. The type species is N. sanctibernardini.
Nipponocypris takayamai^[158]	Sp. nov	Valid	Miyata, Yabumoto & Hirano	Middle Pleistocene	Nogami		A species of Nipponocypris.
Palealbula declivis	Sp. nov	Valid	Schwarzhans	Early Cretaceous (Valanginian)			A member of Albuliformes of uncertain phylogenetic placement.
Palealbula depressidorsalis	Sp. nov	Valid	Schwarzhans	Early Cretaceous (Albian)			A member of Albuliformes of uncertain phylogenetic placement.
Paleoschizothorax^[159]	Gen. et sp. nov	Valid	Yang et al.	Oligocene	Qaidam Basin		A member of the family Cyprinidae related to members of the genus Schizothorax. Genus includes new species P. qaidamensis.
Paleoserranus^[160]	Gen. et sp. nov	Valid	Cantalice, Alvarado-Ortega & Alaniz-Galvan	Paleocene			A member of the family Serranidae. Genus includes new species P. lakamhae.
Parahaplolepis poppaea	Sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Haplolepidae.
Peltopleurus tyrannos^[161]	Sp. nov	Valid	Xu, Ma & Zhao	Middle Triassic (Ladinian)	Falang
Pickeringius^[162]	Gen. et sp. nov	Valid	Choo et al.	Late Devonian	Gogo		An early ray-finned fish. Genus includes new species P. acanthophorus.
Piranhamesodon^[163]	Gen. et sp. nov	Valid	Kölbl-Ebert et al.	Late Jurassic (late Kimmeridgian to early Tithonian)			A member of Pycnodontiformes of uncertain phylogenetic placement. The type species is P. pinnatomus.
Pontosciaena	Gen. et comb. nov	Valid	Bannikov, Schwarzhans & Carnevale	Neogene		Crimean Peninsula? ?	A member of the family Sciaenidae. The type species is "Serranus" acuterostratus Rückert-Ülkümen (1996); genus might also include "Otolithus (Percidarum)" sigmoilinoides Pobedina (1956).
Protalbula websteri	Sp. nov	Valid	Schwarzhans	Late Jurassic (Kimmeridgian)			A member of Albuliformes of uncertain phylogenetic placement.
Protarpon boualii^[164]	Sp. nov	Valid	Khalloufi et al.	Paleocene (Danian)	Oulad Abdoun Basin		A relative of tarpons.
Protoelops	Gen. et comb. nov	Valid	Schwarzhans	Middle Jurassic to Early Cretaceous (Bathonian to Aptian)			A member of the family Elopidae. The type species is "Otolithus (Leptolepidarum)" cuneiformis Frost (1924); genus also includes "Leptolepis" tenuirostris Stinton (1968) and "genus Protacanthopterygiorum" scalpellum Nolf (2004).
Protosciaena kirbyorum^[165]	Sp. nov	Valid	Stringer & Bell	Early Pliocene		United States	A member of the family Sciaenidae.
Pseudogonatodus aurulentum	Sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Gonatodidae.
Pseudohaplolepis	Gen. et sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri of uncertain phylogenetic placement. The type species is P. argentatum.
Pseudotrichiurus	Gen. et sp. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian)	Eutaw		A member of Aulopiformes of uncertain phylogenetic placement. The type species is P. sagax.
Pterothrissus carolinensis^[166]	Sp. nov	Valid	Stringer et al.	Late Cretaceous (early Campanian)	Tar Heel		A relative of the Japanese gissu.
Quasimodichthys^[167]	Gen. et comb. nov	Valid	Paiva & Gallo	Late Jurassic (Oxfordian)	Pastos Bons		A member of Semionotiformes. The type species is "Lepidotus" piauhyensis Roxo & Löfgren (1936)
Rhadinichthys glabrolepis	Sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Rhadinichthyidae.
Rhadinichthys? ornatocephalum	Sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Rhadinichthyidae.
Rhadinichthys? plumosum	Sp. nov	Valid	Elliott	Carboniferous (Bashkirian)	Lower Coal Measures		A member of Actinopteri belonging to the family Rhadinichthyidae.
Sanjuanableps^[168]	Gen. et sp. nov	Valid	Bogan et al.	Late Miocene			A member of the family Anablepidae. Genus includes new species S. calingasta.
Saurichthys spinosa^[169]	Sp. nov	Valid	Wu, Sun & Fang	Middle Triassic (Anisian)	Guanling
Sciades latissimum	Sp. nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco		A species of Sciades.
Sciades peregrinus	Sp. nov	Valid	Aguilera & Marceniuk	Late Miocene	Urumaco		A species of Sciades.
Scopulipiscis^[170]	Gen. et sp. nov	Valid	Latimer & Giles	Late Triassic	Kössen		A relative of Dapedium. Genus includes new species S. saxciput.
Sigmapycnodus	Gen. et sp. nov	Valid	Taverne & Capasso	Late Cretaceous (Cenomanian)			A member of the family Pycnodontidae. The type species is S. giganteus.
Sphaeronchus rundlei	Sp. nov	Valid	Schwarzhans	Late Jurassic (Kimmeridgian)	Kimmeridge Clay		A member of Leptolepidiformes sensu lato of uncertain phylogenetic placement.
Vinctifer ferrusquiai^[171]	Sp. nov	Valid	Cantalice, Alvarado-Ortega & Brito	Late Jurassic (Kimmeridgian)	Sabinal		A member of Aspidorhynchiformes belonging to the family Aspidorhynchidae.
Vox thlotlo	Gen. et sp. nov	Valid	Schwarzhans, Huddleston & Takeuchi	Late Cretaceous (Santonian)	Eutaw		A teleost of uncertain phylogenetic placement. The type species is V. thlotlo.
Wettonius^[172]	Gen. et sp. nov	Valid	Carnevale & Bannikov	Eocene	Monte Bolca Nanjemoy Formation	^[173]	A member of the family Veliferidae. Genus includes new species W. angeloi.
Xenoleptolepis	Gen. et comb. nov	Valid	Schwarzhans	Early Jurassic to Early Cretaceous (Sinemurian to Berriasian)			A member of the family Leptolepididae sensu lato. The type species is Otolithus (incertae sedis) withersi Frost (1926); genus also includes Otolithus (Salmonoidei) oncorhynchoides Weiler (1954).
Yuskaichthys^[174]	Gen. et sp. nov	Valid	Bogan, Agnolin & Scanferla	Eocene	Maíz Gordo		A member of the family Andinichthyidae. The type species is Y. eocenicus.
Zaprora koreana^[175]	Sp. nov	Valid	Nam & Nazarkin	Neogene			A relative of the prowfish.
Zurupleuropholis^[176]	Gen. et 2 sp. nov	Valid	Giordano et al.	Early Cretaceous (Albian)	Lagarcito		A relative of Pleuropholis. Genus includes new species Z. quijadensis and Z. decollavi.

Lobe-finned fishes

Name	Novelty	Status	Authors	Age	Unit	Notes
Celsiodon^[177]	Gen. et sp. nov	Valid	Clack et al.	Devonian (late Famennian)		A lungfish related to Ctenodus. Genus includes new species is C. ahlbergi.
Ceratodus tunuensis^[178]	Sp. nov	Valid	Agnolin et al.	Late Triassic	Fleming Fjord	A lungfish.
Eusthenopteron jenkinsi^[179]	Sp. nov	Valid	Downs et al.	Devonian (Frasnian)	Fram
Mawsonia soba^[180]	Sp. nov	Valid	Brito et al.	Early Cretaceous	Babouri Figuil Basin	A coelacanth.
Ptychoceratodus oldhami^[181]	Sp. nov	Valid	Bhat & Ray	Late Triassic (Carnian)	Tiki	A lungfish. Announced in 2018; the final version of the article naming it was published in 2020.

Notes and References

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