2012 in paleobotany explained

This paleobotany list records new fossil plant taxa that were to be described during the year 2012, as well as notes other significant paleobotany discoveries and events which occurred during 2012.

Ferns and fern allies

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Korallipteris^[4]	Gen. et comb. nov	Valid	Vera & Passalia	Triassic to Miocene		Antarctica ^[5]	A fern of uncertain phylogenetic placement. A new genus for "Gleichenia" argentinica (1962); Additional species were originally placed in Gleichenia, Gleichenites and Microphyllopteris.
Millerocaulis tekelili^[6]	Sp. nov	Valid	Vera	Early Cretaceous (Aptian)	Cerro Negro Formation	Antarctica (Livingston Island)	An Osmundaceae fern. Originally placed in Millerocaulis; Moved to Claytosmunda tekelili in (2017).^[7]
Osmundopsis rafaelii^[8]	Sp. nov	Valid	Escapa & Cúneo	Early Jurassic			A member of Osmundaceae.
Todites cacereii	Sp. nov	Valid	Escapa & Cúneo	Early Jurassic			A member of Osmundaceae.

Bennettitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Anomozamites harrisi^[9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Anomozamites heeri	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Anomozamites villosus^[10]	Sp. nov	Valid	Pott et al.	Probably late Middle Jurassic	Daohugou Beds		A member of Bennettitales. Originally described as a species of Anomozamites Moved to the Wielandiella villosus (2015).^[11] ^[12]
Cycadolepis barnardii	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis karatauensis	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis kirillii	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis kokalensis	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis lata	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis maija	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis multistomata	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis pachydermatica	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis perfecta	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis prynadае	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Cycadolepis schairensis	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Nilssoniopteris chirchilica	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Кokala Suite
Nilssoniopteris localis	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Nilssoniopteris saveljevii	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Tonasha Formation
Pterophyllum chagaense	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Pterophyllum doludenkoae	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Pterophyllum originalum	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Pterophyllum pаrvisegmentatum	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Pterophyllum pinniforme	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Pterophyllum valentinaeа	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Pterophyllum vsevolodii	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Williamsonia antonia	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Williamsonia baranovaeа	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Williamsonia chagaensе	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Williamsonia kokalensis	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Williamsonia vialovaea	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite

"Pteridospermatophytes"

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ctenozamites schweitzerii	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation		A member of Pteridospermatophyta of uncertain affinities.
Eretmonia macloughlinii^[13]	Sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Upper Buckley Formation		A member of Glossopteridales.
Kuvakospermum^[14]	Gen. et sp. nov	Valid	Naugolnykh & Sidorov	Permian (Wordian)			A seed fern belonging to the family Peltaspermaceae. Genus includes new species K. pedatum.
Lidgettoniopsis^[15]	Gen. et sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Beacon Supergroup (upper Buckley Formation)		A seed fern belonging to the group Dictyopteridiales and the family Eretmoniaceae. Genus includes new species L. ramulus.
Scutum leiophyllum	Sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Beacon Supergroup (upper Buckley Formation)		A member of Glossopteridales belonging to the family Dictyopteridiaceae.
Vetlugospermum^[16]	Gen. et sp. nov	Valid	Naugolnykh	Early Triassic			A seed fern belonging to the group Peltaspermales. Genus includes new species V. rombicum.

Ginkgoales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Baiera kazachstanica	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite
Baieroxylon rocablanquense^[17]	Sp. nov	Valid	Gnaedinger	Early Jurassic	Roca Blanca Formation
Ginkgo cranei^[18]	Sp. nov	Valid	Zhou, Quan, & Liu	upper Paleocene	Sentinel Butte Formation		An extinct species in the modern genus Ginkgo
Ginkgo gomolitzkyana^[19]	Sp. nov	Valid	Nosova	Middle Jurassic	Angren Formation		A member of Ginkgoales.
Ginkgo huolinhensis^[20]	Sp. nov	Valid	Dong & Sun	Early Cretaceous	Huolinhe Formation
Ginkgoites barnardianus	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Кokala Suite
Ginkgoites irdalicus	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic	Ergozi Suite
Ginkgoites kelendensis	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Tonasha Formation
Ginkgoites ketovianus	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Bathonian)	Sarydiirmeny Suite
Leptotoma asiatica	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation
Pseudotorellia karadiirmenica	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation

Conifers

Cheirolepidiaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Duartenia^[22]	Gen. et sp. nov	Valid	Mohr et al.	Early Cretaceous	Crato Formation		A possible member of Cheirolepidiaceae. The type species is Duartenia araripensis.
Glenrosa falcata^[23]	Sp. nov	Valid	Gomez, Ewin & Daviero-Gomez	Early Cretaceous (Barremian)	Calizas de La Huergina Formation		A conifer.

Cupressaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Calocedrus huashanensis^[24]	Sp. nov	Valid	Shi, Zhou, & Xie	Oligocene	Ningming Formation		A Calocedrus species
Fokienia shengxianensis^[25]	Sp. nov	Valid	He, Sun & Liu	Late Miocene	Shengxian Formation Xiaolongtan Formation		Originally placed in Fokienia. Moved to Calocedrus shengxianensis in (2015).^[26]

Pinaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Eathiestrobus^[27]	Gen. et sp. nov	Valid	Rothwell et al.	Late Jurassic (Kimmeridgian)	Kimmeridge Clay Formation		A member of Pinaceae. The type species is Eathiestrobus mackenziei.
Picea burtonii^[28]	Sp. nov	Valid	Klymiuk & Stockey	Early Cretaceous (Valanginian)			A spruce species seed cone.
Pinus yorkshirensis^[29]	Sp. nov	Valid	Ryberg et al.	Early Cretaceous (Hauterivian-Barremian)	Speeton Clay Formation		A pine.

Taxaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Marskea kazachstanica	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Kokala Suite
Torreya kazachstanica	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Kokala Suite

Other conifers

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Elatocladus bulakus	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Kokala Suite		A conifer of uncertain affinities.
Plyophyllioxylon^[31]	Gen. et sp. nov	Valid	Feng et al.	Permian (Roadian/Wordian)	Lower Shihhotse Formation		A conifer of uncertain phylogenetic placement. Genus includes new species P. hulstaiense.

Other gymnosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Nilssonia prosvirjakovaea	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation		A cycad.
Rehezamites^[32]	Gen. et sp. nov	Valid	Pott et al.	Early Cretaceous (Aptian)	Yixian Formation		A seed plant of uncertain phylogenetic placement. Genus includes new species R. anisolobus.

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Archaeampelos senonica^[33]	Sp. nov	Valid	Alekseev	Late Cretaceous
Baasia^[34]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation		A member of the family Celastraceae. Genus includes new species B. armendarisense.
Bertilanthus^[35]	Gen. et sp. nov	Valid	Friis & Pedersen	Late Cretaceous			An asterid. Genus includes new species B. scanicus.
Camptodromites sibiricus	Sp. nov	Valid	Alekseev	Late Cretaceous
Celastrinites lanceolatus	Sp. nov	Valid	Alekseev	Late Cretaceous
Chaneya hainanensis^[36]	Sp. nov	Valid	Feng & Jin	Eocene	Changchang Basin		A fossil fruit.
Cissites basicordatus	Sp. nov	Valid	Alekseev	Late Cretaceous
Cissus lombardii^[37]	Sp. nov	Valid	Manchester, Chen & Lott	Early Oligocene	Mancora Formation		A species of Cissus.
Eucalyptus caldericola^[38]	Sp. nov	Valid	Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)			A species of Eucalyptus.
Eucalyptus frenguelliana	Sp. nov	Valid	Gandolfo & Zamaloa in Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)			A species of Eucalyptus.
Eucalyptus lynchiae	Sp. nov	Valid	Gandolfo & Hermsen in Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)			A species of Eucalyptus.
Fulleroxylon	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation		A member of the family Myrtaceae. Genus includes new species F. armendarisense.
Juglandiphyllites microdentatus	Sp. nov	Valid	Alekseev	Late Cretaceous
Laurinoxylon stichkai^[39]	Sp. nov	Valid	Boonchai & Manchester	Early Eocene	Bridger Formation		A member of Lauraceae described on the basis of fossil wood.
Laurophyllum calicarioides	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Litsea.
Laurophyllum lacustris	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum maarensis^[40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Cryptocarya.
Laurophyllum microphyllum	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Cryptocarya.
Laurophyllum otagoensis	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum sylvestris	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum taieriensis	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Cryptocarya.
Laurophyllum vulcanicola	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Beilschmiedia.
Laurophyllum waipiata	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae with affinities to members of the genus Cryptocarya.
Litseopsis nova-zelandiae	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene			A member of Lauraceae.
Mahonia grimmii^[41]	Sp. nov	Valid	Güner & Denk	Miocene	Eskihisar Formation		A species of Mahonia.
Mahonia somaensis	Sp. nov	Valid	Güner & Denk	Miocene	Soma Formation		A species of Mahonia.
Morinda chinensis^[42]	Sp. nov	Valid	Shi et al.	Eocene	Changchang Formation		A species of Morinda.
Newtonia mushensis^[43]	Sp. nov	Valid	Pan et al.	Early Miocene			A species of Newtonia.
Palaeophytocrene hammenii^[44]	Sp. nov	Valid	Stull in Stull et al.	Middle-late Paleocene	Bogotá Formation		A member of Icacinaceae belonging to the tribe Phytocreneae.
Palaeophytocrene piggae	Sp. nov	Valid	Stull in Stull et al.	Late Paleocene	Sentinel Butte Formation		A member of Icacinaceae belonging to the tribe Phytocreneae.
Pygmaeoxylon	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation		A magnoliid of uncertain phylogenetic placement. Genus includes new species P. paucipora.
Pyrenacantha austroamericana	Sp. nov	Valid	Stull in Stull et al.	Late early Oligocene			A species of Pyrenacantha.
Schenkeriphyllum glanduliferum^[45]	Gen. et sp. nov	Valid	Mohr, Coiffard & Bernardes-de-Oliveira	Early Cretaceous (Aptian)	Crato Formation		Fossil magnolialean angiosperm.
Wilsonoxylon	Gen. et sp. nov	Valid	Boonchai & Manchester	Early Eocene	Bridger Formation		A member of Canellaceae described on the basis of fossil wood. Genus includes new species W. edenense.

Notes and References

1. Ignatov . M. S. . Karasev . E. V. . Sinitsa . S. M. . Maslova . E. V. . New Bryokhutuliinia species (Bryophyta) with sporophytes from the Upper Jurassic of Transbaikalia. . Arctoa . 2013 . 22 . 69–78. 10.15298/arctoa.22.10 . free .
2. Michael T. Dunn . Prescott Atkinson . James Lacefield . Michael Rischbieter . 2012 . Winslowia tuscumbiana gen. et sp. nov. (Chaloneriaceae): A Cormose, Heterosporous, Ligulate Lycopsid Reconstructed from the Inside Out from the Pride Mountain Formation (Late Mississippian/Serpukhovian) of Northern Alabama . International Journal of Plant Sciences . 173 . 1 . 96–111 . 10.1086/662679 . 85874193 .
3. Cai-Qing Guo . Dianne Edwards . Peng-Cheng Wu . Jeffrey G. Duckett . Francis M. Hueber . Cheng-Sen Li . 2012 . Riccardiothallus devonicus gen. et sp. nov., the earliest simple thalloid liverwort from the Lower Devonian of Yunnan, China . Review of Palaeobotany and Palynology . 176–177 . 35–40 . 10.1016/j.revpalbo.2012.03.012 . 2012RPaPa.176...35G .
4. Ezequiel I. Vera . Mauro G. Passalia . 2012 . Korallipteris, a new genus for Mesozoic Gleichenia-like fern fronds . Geobios . 45 . 4 . 421–428 . 10.1016/j.geobios.2011.09.002 . 2012Geobi..45..421V . 11336/68155 . free .
5. John G. Conran . Joseph A. Jackson . Daphne E. Lee . Elizabeth M. Kennedy . 2017 . Gleichenia-like Korallipteris alineae sp. nov. macrofossils (Polypodiophyta) from the Miocene Landslip Hill silcrete, New Zealand . New Zealand Journal of Botany . 55 . 3 . 258–275 . 10.1080/0028825X.2017.1317278 . 2017NZJB...55..258C . 91077403 .
6. Ezequiel I. Vera . 2012 . Millerocaulis tekelili sp. nov., a new species of osmundalean fern from the Aptian Cerro Negro Formation (Antarctica) . Alcheringa: An Australasian Journal of Palaeontology . 36 . 1 . 35–45 . 10.1080/03115518.2011.576541 . 2012Alch...36...35V . 129810694 .
7. Benjamin Bomfleur . Guido W. Grimm . Stephen McLoughlin . 2017 . The fossil Osmundales (Royal Ferns)—a phylogenetic network analysis, revised taxonomy, and evolutionary classification of anatomically preserved trunks and rhizomes . PeerJ . 5 . e3433 . 10.7717/peerj.3433 . 28713650 . 5508817 . free .
8. Ignacio H. Escapa . N. Rubén Cúneo . 2012 . Fertile Osmundaceae from the Early Jurassic of Patagonia, Argentina . International Journal of Plant Sciences . 173 . 1 . 54–66 . 10.1086/662652 . 85319731 . 11336/194430 . free .
9. Book: Anna I. Kiritchkova . Natalya V. Nosova . 2012 . Jurassic continental deposits of the Middle-Caspian Basin. Part 2: Facies, taphonomy, interregional correlations, flora (Pinophyta: Pteridospermae, Cycadales, Bennettitales, Ginkgoales, Czekanowskiales, Coniferales) . 1–338 . VNIGRl . 978-5-88953-147-0 .
10. Christian Pott . Stephen McLoughlin . Shunqing Wu . Else Marie Friis . 2012 . Trichomes on the leaves of Anomozamites villosus sp. nov. (Bennettitales) from the Daohugou beds (Middle Jurassic), Inner Mongolia, China: Mechanical defence against herbivorous arthropods . Review of Palaeobotany and Palynology . 169 . 48–60 . 10.1016/j.revpalbo.2011.10.005 . 2012RPaPa.169...48P .
11. Christian Pott . Wang Xiaoli . Zheng Xiaoting . 2015 . Wielandiella villosa comb. nov. from the Middle Jurassic of Daohugou, China: More evidence for divaricate plant architecture in Williamsoniceae . Botanica Pacifica . 4 . 2 . 137–148 . 10.17581/bp.2015.04215 . free .
12. Christian Pott . Baoyu Jiang . 2017 . Plant remains from the Middle–Late Jurassic Daohugou site of the Yanliao Biota in Inner Mongolia, China . Acta Palaeobotanica . 57 . 2 . 185–222 . 10.1515/acpa-2017-0012 . 62827595 . free .
13. Patricia E. Ryberg . Edith L. Taylor . Thomas N. Taylor . 2012 . The First Permineralized Microsporophyll of the Glossopteridales: Eretmonia macloughlinii sp. nov. . International Journal of Plant Sciences . 173 . 7 . 812–822 . 10.1086/666667 . 1808/13676 . 54003097 . free .
14. Book: S.V. Naugolnykh . A.A. Sidorov . 2012 . A new representative of gymnosperms of the order Peltaspermales from the Novy Kuvak locality (Kazanian stage; Samara region, Russia) . S.V. Naugolnykh . Palaeontology and evolution of the biodiversity in the Earth history (in museum context). Collection of scientific articles . GEOS . 71–78 . 978-5-89118-575-3 .
15. Patricia E. Ryberg . Edith L. Taylor . Thomas N. Taylor . 2012 . Antarctic glossopterid diversity on a local scale: The presence of multiple megasporophyll genera, Upper Permian, Mt. Achernar, Transantarctic Mountains, Antarctica . American Journal of Botany . 99 . 9 . 1531–1540 . 10.3732/ajb.1200142 . 22922397 .
16. Serge V. Naugolnykh . 2012 . Vetlugospermum and Vetlugospermaceae: A new genus and family of peltasperms from the Lower Triassic of Moscow syneclise (Russia) . Geobios . 45 . 5 . 451–462 . 10.1016/j.geobios.2011.10.009 . 2012Geobi..45..451N .
17. Silvia Gnaedinger . 2012 . Ginkgoalean woods from the Jurassic of Argentina: Taxonomic considerations and palaeogeographical distribution . Geobios . 45 . 2 . 187–198 . 10.1016/j.geobios.2011.01.007 . 2012Geobi..45..187G . 11336/25674 . free .
18. Zhou . Z. . Quan . C. . Liu . Y-S . 86289858 . 2012 . Tertiary Ginkgo ovulate organs with associated leaves from North Dakota, U.S.A., and their evolutionary significance . International Journal of Plant Sciences . 173 . 1 . 67–80 . 10.1086/662651.
19. Nosova . Natalya . 2012 . Женские репродуктивные структуры Ginkgo gomolitzkyana N. Nosova, sp. nov. из среднеюрских отложений Ангрена (Узбекистан) / Female reproductive structures of Ginkgo gomolitzkyana N. Nosova, sp. nov. from the Middle Jurassic of Angren (Uzbekistan) . Paleobotanika . 3 . 62–91 . 10.31111/palaeobotany/2012.3.62 .
20. Man Dong . Ge Sun . 2012 . Ginkgo huolinhensis sp. nov. from the Lower Cretaceous of Huolinhe Coal Field, Inner Mongolia, China . Acta Geologica Sinica (English Edition) . 86 . 1 . 11–19 . 10.1111/j.1755-6724.2012.00607.x . 2012AcGlS..86...11D . 129029365 .
21. Mary E. Dettmann . H. Trevor Clifford . Mark Peters . 2012 . Emwadea microcarpa gen. et sp. nov.—anatomically preserved araucarian seed cones from the Winton Formation (late Albian), western Queensland, Australia . Alcheringa: An Australasian Journal of Palaeontology . 36 . 2 . 217–237 . 10.1080/03115518.2012.622155 . 2012Alch...36..217D . 129171237 .
22. Mohr . B.A.R. . Schultka . S. . Süss . H. . Bernardes-de-Oliveira . M.E.C. . 2012 . A new drought resistant gymnosperm taxon Duartenia araripensis gen. nov. et sp. nov. (Cheirolepidiaceae?) from the Early Cretaceous of Northern Gondwana. . Palaeontographica Abteilung B . 289 . 1–3 . 1–25 . 2013-03-28 . https://web.archive.org/web/20130426013110/http://www.schweizerbart.de/papers/palb/detail/289/79289/A_new_drought_resistant_gymnosperm_taxon_Duartenia . 2013-04-26 . dead . 10.1127/palb/289/2012/1 . 2012PalAB.289....1M .
23. Gomez . Bernard . Ewin . Timothy A.M. . Daviero-Gomez . Véronique . 2012 . The conifer Glenrosa falcata sp. nov. from the Lower Cretaceous of Spain and its palaeoecology . Review of Palaeobotany and Palynology . 172 . 21–32 . 10.1016/j.revpalbo.2012.01.009 . 2012RPaPa.172...21G .
24. Shi . G. . Zhou . Z. . Xie . Z. . 2012 . A new Oligocene Calocedrus from south China and its implications for transpacific floristic exchanges. . American Journal of Botany . 99 . 1 . 108–120 . 10.3732/ajb.1100331 . 22223689.
25. Wenlong He . Bainian Sun . Yu-Sheng (Christopher) Liu . 2012 . Fokienia shengxianensis sp. nov. (Cupressaceae) from the late Miocene of eastern China and its paleoecological implications . Review of Palaeobotany and Palynology . 176–177 . 24–34 . 10.1016/j.revpalbo.2012.03.013 . 2012RPaPa.176...24H .
26. Jian-Wei Zhang . Jian Huang . Ashalata D'Rozario . Jonathan M. Adams . Zhe-Kun Zhou . 2015 . Calocedrus shengxianensis, a late Miocene relative of C. macrolepis (Cupressaceae) from South China: Implications for paleoclimate and evolution of the genus . Review of Palaeobotany and Palynology . 222 . 1–15 . 10.1016/j.revpalbo.2015.07.004 . 2015RPaPa.222....1Z .
27. Gar W. Rothwell . Gene Mapes . Ruth A. Stockey . Jason Hilton . 2012 . The seed cone Eathiestrobus gen. nov.: Fossil evidence for a Jurassic origin of Pinaceae . American Journal of Botany . 99 . 4 . 708–720 . 10.3732/ajb.1100595 . 22491001.
28. Ashley A. Klymiuk . Ruth A. Stockey . 2012 . A Lower Cretaceous (Valanginian) seed cone provides the earliest fossil record for Picea (Pinaceae) . American Journal of Botany . 99 . 6 . 1069–1082 . 10.3732/ajb.1100568 . 22623610 . free .
29. Patricia E. Ryberg . Gar W. Rothwell . Ruth A. Stockey . Jason Hilton . Gene Mapes . James B. Riding . 2012 . Reconsidering Relationships among Stem and Crown Group Pinaceae: Oldest Record of the Genus Pinus from the Early Cretaceous of Yorkshire, United Kingdom . International Journal of Plant Sciences . 173 . 8 . 917–932 . 10.1086/667228 . 85402168 .
30. Peter Wilf . 2012 . Rainforest conifers of Eocene Patagonia: Attached cones and foliage of the extant Southeast Asian and Australasian genus Dacrycarpus (Podocarpaceae) . American Journal of Botany . 99 . 3 . 562–584 . 10.3732/ajb.1100367 . 22334450 .
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