2009 in arthropod paleontology explained

This list of fossil arthropods described in 2009 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods that have been described during the year 2009, as well as other significant discoveries and events related to arthropod paleontology that occurred.

Crustaceans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Chabardella^[3]	Gen. et sp. nov	Valid	Racheboeuf, Schram & Vidal	Late Carboniferous Stephanian	Montceau-les-Mines		A mantis shrimp, type species is C. spinosa
Sottyella	Gen. et sp. nov	Valid	Racheboeuf, Schram & Vidal	Late Carboniferous Stephanian	Montceau-les-Mines		A phreatoicidean isopod, type species is S. montcellensis

Insects

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Agulla mineralensis^[4]	sp nov	Valid	Engel	Late Barstovian	Stewart Valley Group		only described Neogene snakefly fossil
Allorapisma^[5]	gen et sp nov	Valid	Makarkin & Archibald	Ypresian	Tom Thumb Tuff member Klondike Mountain Formation
Apis (Cascapis) nearctica^[6]	sp nov	Valid	Engel, Hinojosa-Diaz, & Rasnitsyn	Middle Miocene		United States	The first fossil Apis from North America.
Aspidopleura^[7]	gen et sp nov	Valid	Gibson	Eocene	Baltic amber		A Eupelmid wasp
Brevivulva	gen et sp nov	Valid	Gibson	Eocene	Baltic amber		A Eupelmid wasp
Cimbrophlebia brooksi^[8]	sp nov	Valid	Archibald	Ypresian	Tom Thumb Tuff member Klondike Mountain Formation		A Cimbrophlebiid scorpionfly
Cimbrophlebia flabelliformis	sp nov	Valid	Archibald	Ypresian	McAbee Fossil Beds		A mecopteran
Cimbrophlebia leahyi	sp nov	Valid	Archibald	Ypresian	McAbee Fossil Beds		A mecopteran
Cimbrophlebia westae	sp nov	Valid	Archibald	Ypresian	Tom Thumb Tuff member Klondike Mountain Formation		A mecopteran
Ctenobethylus oblongiceps^[9]	comb nov	Jr synonym	(Wheeler, 1915)	Lutetian	Baltic amber		Fossil Dolichoderine ant. Moved from Iridomyrmex oblongiceps to Eldermyrmex oblongiceps in 2011.
Denaeaspis^[10]	gen et sp nov	valid	Chaboo & Engel	Eocene Bridgerian	Green River Formation Parachute Creek Member		An Imatidiini tribe Cassidine tortoise beetle. The type species is D. chelonopsis
Eosacantha	gen et sp nov	valid	Chaboo & Engel	Eocene Bridgerian	Green River Formation Parachute Creek Member		An Imatidiini tribe Cassidine tortoise beetle. The type species is E. delocranioides
Gesomyrmex germanicus^[11]	sp nov	Valid	Dlussky, Wappler, & Wedmann	Lutetian	Eckfeld Maar		A Formicinae ant
Gesomyrmex pulcher	sp nov	Valid	Dlussky, Wappler, & Wedmann	Lutetian	Messel Formation		A Formicinae ant
Leptofoenus pittfieldae^[12]	sp nov	Valid	Engel	Burdigalian	Dominican amber		A Pteromalid wasp
Metapelma archetypon	sp nov	Valid	Gibson	Eocene	Baltic amber		A Eupelmid wasp
Neanaperiallus	gen et sp nov	Valid	Gibson	Eocene	Baltic amber		A Eupelmid wasp
Nesagapostemon^[13]	gen et sp nov	Valid	Engel	Burdigalian	Dominican amber		A Halictid bee
Nymphes georgei^[14]	sp nov	Valid	Archibald, Makarkin, & Ansorge	Ypresian	Tom Thumb Tuff member Klondike Mountain Formation		A Nymphid neuropteran
Oligochlora semirugosa	sp nov	Valid	Engel	Burdigalian	Dominican amber		A Halictid bee
Paleohabropoda^[15]	Gen et sp nov	valid	Michez & Ramont	Thanetian	Menat Formation		An anthophorine bee
Paratrechina pygmaea^[16]	Comb nov	Jr synonym	(Mayr)	Middle Eocene	European amber		A formicine ant, new combination for Prenolepis pygmaea; jr synonym of Nylanderia pygmaea
Principiala rudgwickensis^[17]	sp nov	valid	Jepson, Makarkin, & Jarzembowski	Barremian	Upper Weald Clay		An Ithonidae lacewing, a second species of Principiala
Proceratium eocenicum^[18]	Sp nov	Valid	Dlussky	Late Eocene	Baltic amber		A ponerine ant
Pronymphes hoffeinsorum	Sp nov	Valid	Archibald, Makarkin, Ansorge	Priabonian	Baltic amber		A Nymphid neuropteran
Protomyrmica^[19]	Gen et sp nov	valid	Dlussky & Radchenko	Priabonian	Baltic amber		A myrmicine ant
Termitaradus mitnicki^[20]	sp nov	Valid	Engel	Burdigalian	Dominican amber		A termite bug
Tetraponera groehni	sp nov	Valid	Dlussky	Pribonian	Baltic amber		A Pseudomyrmecine ant

Trilobites

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Alperillaenus^[21]	Gen. et sp. nov	Valid	Pour, Popov & Vinogradova	Ordovician (Darriwilian)			A member of the family Illaenidae. Genus includes new species A. intermedius.
Ammagnostus antarcticus	Sp. nov	Valid	Bentley, Jago & Cooper	Cambrian	Goyder Formation^[22] Spurs Formation	Antarctica
Asaphellus stubbsi^[23]	Sp. nov	Valid	Fortey	Ordovician (Tremadocian)	Fezouata Formation
Bhargavia^[24]	Gen. et sp. et comb. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Ellipsocephalidae. The type species is B. prakritika; genus also includes "Syspacephalus" obscurus Palmer and Halley (1979).
Bienvillia chaya^[25]	Sp. nov	Valid	Waisfeld & Vaccari	Ordovician (Floian)	Suri Formation
Catillicephalina^[26]	Gen. et comb. nov	Valid	Bentley, Jago & Cooper	Cambrian	Spurs Formation	Antarctica	A new genus for "Catillicephala" glasgowensis Jago & Cooper (2005).
Damiraspis	Gen. et sp. nov	Valid	Pour, Popov & Vinogradova	Ordovician (Darriwilian)			An asaphid trilobite. Genus includes new species D. margiana.
Degamella lingulata^[27]	Sp. nov	Valid	Zhou & Zhou	Ordovician	Pagoda Formation		A member of the family Cyclopygidae.
Dividuagnostus tortelloi	Sp. nov	Valid	Waisfeld & Vaccari	Ordovician (Floian)	Suri Formation
Doremataspis contracta	Sp. nov	Valid	Bentley, Jago & Cooper	Cambrian	Spurs Formation	Antarctica
Farasaphus	Gen. et sp. nov	Valid	Pour, Popov & Vinogradova	Ordovician (Darriwilian)			An asaphid trilobite. Genus includes new species F. singularis.
Gunnia smithi	Sp. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Ptychopariidae.
Haydenaspis	Gen. et sp. nov	Valid	Peng et al.	Cambrian Stage 4	Parahio Formation		A member of the family Dinesidae. The type species is H. parvatya.
Himalisania	Gen. et comb. nov	Valid	Peng et al.	Cambrian (Guzhangian)	Karsha Formation Kurgiakh Formation	?	A member of the family Lisaniidae. The type species is "Eoshengia" sudani Jell & Hughes (1997).
Hypermecaspis edselbrussai	Sp. nov	Valid	Waisfeld & Vaccari	Ordovician (Floian)	Suri Formation
Jamrogia	Gen. et sp. nov	Valid	Bentley, Jago & Cooper	Cambrian	Spurs Formation	Antarctica	A member of the new family Jamrogiidae. Genus includes new species J. jamrogensis.
Kaotaia prachina	Sp. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Ptychopariidae.
Koldinia odelli	Sp. nov	Valid	Peng et al.	Cambrian (Guzhangian)	Karsha Formation		Possibly a member of the family Anomocaridae.
Litzicurus^[28]	Gen. et sp. et comb. nov	Valid	Adrain, McAdams & Westrop	Early Ordovician			Possibly a member of Hystricuridae. Genus includes new species L. shawi, as well as "Pseudohystricurus" orbus Ross (1953).
Microparia (Quadratapyge) obsoleta	Sp. nov	Valid	Zhou & Zhou	Ordovician	Pagoda Formation		A member of the family Cyclopygidae.
Monanocephalus liquani	Sp. nov	Valid	Peng et al.	Wuliuan	Hsuchuang Formation		A member of the family Ptychopariidae.
Nileus jafari^[29]	Sp. nov	Valid	Pour & Turvey	Ordovician (Dapingian to early Darriwilian)	Shirgesht Formation		A member of the family Nileidae.
Proasaphiscus simoni	Sp. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Proasaphiscidae.
Probowmania bhatti	Sp. nov	Valid	Peng et al.	Cambrian Stage 4	Parahio Formation		A member of the family Ptychopariidae.
Prozacanthoides lahiri	Sp. nov	Valid	Peng et al.	Cambrian Stage 4	Parahio Formation		A member of the family Zacanthoididae.
Psalikilus hestoni	Sp. nov	Valid	Adrain, McAdams & Westrop	Early Ordovician
Sudanomocarina sinindica	Sp. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Proasaphiscidae.
Torifera jelli	Sp. nov	Valid	Peng et al.	Cambrian (Guzhangian)	Karsha Formation Kurgiakh Formation		A member of the family Alsataspididae.
Warburtonella^[30]	Gen. et sp. nov	Valid	Sun & Jago	Cambrian (Furongian)	Warburton Basin		A polymerid trilobite. Genus includes new species W. coongiensis.
Xingrenaspis parthiva	Sp. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Ptychopariidae.
Xingrenaspis shyamalae	Sp. nov	Valid	Peng et al.	Wuliuan	Parahio Formation		A member of the family Ptychopariidae.

Notes and References

1. Kühl . G . Briggs . DE . Rust . J . Gabriele Kühl, Derek E. G. Briggs, Jes Rust . A Great-Appendage Arthropod with a Radial Mouth from the Lower Devonian Hunsrück Slate, Germany . Science . 323 . 5915 . 771 - 3 . February 2009 . 0036-8075 . 19197061 . 10.1126/science.1166586 . 2009Sci...323..771K.
2. Selden . P.A. . Penney . D. . 2009 . A fossil spider (Araneae: Pisauridae) of Eocene age from Horsefly, British Columbia, Canada . Contributions to Natural History . 12 . 1269–1282 .
3. Racheboeuf . Patrick R. . Schram . Frederick R. . Vidal . Muriel . 2009 . New Malacostracan Crustacea from the Carboniferous (Stephanian) Lagerstätte of Montceau-Les-Mines, France . Journal of Paleontology . 83 . 4 . 624–629 . 0022-3360.
4. Engel . M. S. . 2009 . A Miocene snakefly from Stewart Valley, Nevada (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science . 112 . 4 . 211–214 . 10.1660/062.112.0409 .
5. Makarkin . V.N. . Archibald, S.B. . 2009 . A new genus and first Cenozoic fossil record of moth lacewings (Neuroptera: Ithonidae) from the Early Eocene of North America . Zootaxa . 2063 . 55–63 . 24 March 2010 . 10.11646/zootaxa.2063.1.3 .
6. Michael S. Engel, I. A. Hinojosa-Diaz & A. P. Rasnitsyn . 2009 . A honey bee from the Miocene of Nevada and the biogeography of Apis (Hymenoptera: Apidae: Apini) . . 60 . 3 . 23–38 . 4 September 2010 .
7. Gibson . G.A.P. . 2009 . Description of three new genera and four new species of Neanastatinae (Hymenoptera, Eupelmidae) from Baltic amber, with discussion of their relationships to extant taxa . ZooKeys . 20 . 10.3897/zookeys.20.161 . 24 March 2010 . 175–214 . https://web.archive.org/web/20100922071426/http://pensoftonline.net/zookeys/index.php/journal/article/view/161 . 22 September 2010 . dead . dmy-all . free .
8. Archibald . S.B. . 2009 . New Cimbrophlebiidae (Insecta: Mecoptera) from the Early Eocene at McAbee, British Columbia, Canada and Republic, Washington, USA . Zootaxa . 2249 . 51–62 . 24 March 2010 . 10.11646/zootaxa.2249.1.5 .
9. Heterick . B. E. . Shattuck . S. . 2011 . Revision of the ant genus Iridomyrmex (Hymenoptera: Formicidae) . Zootaxa . 2845 . 1–174.
10. Chaboo . C. S. . Engel . M. S. . Eocene tortoise beetles from the Green River Formation in Colorado, U.S.A. (Coleoptera: Chrysomelidae: Cassidinae) . Systematic Entomology . 2009 . 34 . 202–209 . 10.1111/j.1365-3113.2008.00456.x . 2.
11. Dlussky . GM . Wappler . T . Wedmann . S . 56253714 . 2009 . Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities . Zootaxa . 2031 . 1–20. 10.11646/zootaxa.2031.1.1 .
12. Engel . M.S. . 2005 . The first fossil leptofoenine wasp (Hymenoptera, Pteromalidae): A new species of Leptofoenusin Miocene amber from the Dominican Republic . ZooKeys . 13 . 57–66. 10.3897/zookeys.13.159. free . 1808/5595 . free .
13. Engel . M.S. . 2009 . Two new halictine bees in Miocene Amber from the Dominican Republic (Hymenoptera, Halictidae) . ZooKeys . 29 . 1–12 . 10.3897/zookeys.29.257. free . 1808/5625 . free .
14. Archibald . S.B. . Makarkin,V.N.. Ansorge, J. . 2009 . New fossil species of Nymphidae (Neuroptera) from the Eocene of North America and Europe . Zootaxa . 2157 . 59–68 . 10.11646/zootaxa.2157.1.4 .
15. Michez . D. . Ramont . P. . 2009 . Taxonomy; New fossil evidence of the early diversification of bees: Paleohabropoda oudardi from the French Paleocene (Hymenoptera, Apidae, Anthophorini) . Zoologica Scripta . 38 . 2 . 171–181 . 10.1111/j.1463-6409.2008.00362.x.
16. LaPolla . J.S. . Dlussky . G.M. . 2010 . Review of fossil Prenolepis genus-group species (Hymenoptera: Formicidae) . Proceedings of the Entomological Society of Washington . 112 . 2 . 258–273 . 10.4289/0013-8797-112.2.258.
17. Jepson . JE . Makarkin . VN . Jarzembowski . E . 2009 . New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden supergroup of Southern England . Cretaceous Research . 30 . 5 . 1325–1338 . 10.1016/j.cretres.2009.07.012.
18. Dlussky . G. M. . 2009 . The ant subfamilies Ponerinae, Cerapachyinae, and Pseudomyrmecinae (Hymenoptera, Formicidae) in the Late Eocene Ambers of Europe . Paleontological Journal . 43 . 9 . 1043–1086 . 10.1134/S0031030109090068 .
19. Dlussky . G. M. . Radchenko . A. G. . 2009 . Two new primitive ant genera from the late Eocene European ambers. . Acta Palaeontologica Polonica . 54 . 3 . 435–441 . 10.4202/app.2008.0092. free .
20. Engel . M.S. . 2009 . A new termite bug in Miocene amber from the Dominican Republic (Hemiptera, Termitaphididae) . ZooKeys . 25 . 61–68. 10.3897/zookeys.25.267. free . 1808/5559 . free .
21. Mansoureh Ghobadi Pour . Leonid E. Popov . Elena V. Vinogradova . Middle Ordovician (late Darriwilian) trilobites from the northern Betpak-Dala Desert, central Kazakhstan . Memoirs of the Association of Australasian Palaeontologists . 37 . 327–349 . 2009 . 0810-8889 .
22. Patrick M. Smith . John R. Paterson . Glenn A. Brock . 2018 . Trilobites and agnostids from the Goyder Formation (Cambrian Series 3, Guzhangian; Mindyallan), Amadeus Basin, central Australia . Zootaxa . 4396 . 1 . 1–67 . 10.11646/zootaxa.4396.1.1 . 29690342 .
23. Richard A. Fortey . A new giant asaphid trilobite from the Lower Ordovician of Morocco . Memoirs of the Association of Australasian Palaeontologists . 37 . 9–16 . 2009 . 0810-8889 .
24. Shanchi Peng . Nigel C. Hughes . Noel A. Heim . Bryan K. Sell . Xuejian Zhu . Paul M. Myrow . Suraj K. Parcha . Cambrian trilobites from the Parahio and Zanskar Valleys, Indian Himalaya . Journal of Paleontology. 71 . 1–95 . 2009 . 10.1666/08-129.1 .
25. Beatriz G. Waisfeld . N. Emilio Vaccari . Trilobites from the Suri Formation (Lower Ordovician; Floian), Famatina Belt, Argentina . Memoirs of the Association of Australasian Palaeontologists . 37 . 407–426 . 2009 . 0810-8889 .
26. Christopher J. Bentley . James B. Jago . Roger A. Cooper . An Acmarhachis typicalis zone trilobite fauna from the Cambrian of northern Victoria Land, Antarctica . Memoirs of the Association of Australasian Palaeontologists . 37 . 165–197 . 2009 . 0810-8889 .
27. Zhiyi Zhou . Zhiqiang Zhou . Ordovician cyclopygid trilobites from the Pagoda Formation of southwestern Shaanxi, China . Memoirs of the Association of Australasian Palaeontologists . 37 . 87–101 . 2009 . 0810-8889 .
28. Jonathan M. Adrain . Neo E.B. McAdams . Stephen R. Westrop . Trilobite biostratigraphy and revised bases of the Tulean and Blackhillsian stages of the Ibexian Series, Lower Ordovician, western United States . Memoirs of the Association of Australasian Palaeontologists . 37 . 541–610 . 2009 . 0810-8889 .
29. Mansoureh Ghobadi Pour . Samuel T. Turvey . Revision of some Lower to Middle Ordovician leiostegiid and associated trilobites from Iran and China . Memoirs of the Association of Australasian Palaeontologists . 37 . 463–480 . 2009 . 0810-8889 .
30. Xiaowen Sun . James B. Jago . An Iverian (Furongian, Cambrian) trilobite faunule from the subsurface Warburton Basin, South Australia . Memoirs of the Association of Australasian Palaeontologists . 37 . 1–8 . 2009 . 0810-8889 .